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Kakui Keiichi

Faculty of Science Biological Sciences BiodiversityLecturer

タナイス目(Tanaidacea)という小型水生甲殻類を主な研究対象にしています.系統分類学的研究を主軸に,性表現,生殖様式,生活様式,形態,行動,寄生生物などについても研究を進めています.タナイス以外についても,指導学生や共同研究者の協力もいただきながら少しずつ研究を進めています.

Researcher basic information

■ Degree
  • Ph. D, Hokkaido University
■ URL
researchmap URLホームページURL■ Various IDs
Researcher number
  • 70723360
ORCID IDJ-Global ID■ Research Keywords and Fields
Research Keyword
  • Tanaidacea
  • Nicothoidae
  • Crustacea
  • Pycnogonida
  • Nectonema
  • Taxonomy
  • Phylogeny
  • 寄生
  • 性表現
  • 生殖様式
  • 雌雄同体
  • 性転換
Research Field
  • Life Science, Biodiversity and systematics, Phylogeny and Taxonomy of Crustaceans
■ Educational Organization

Career

■ Career
Committee Memberships
  • Jan. 2026 - Present
    日本動物分類学会, 庶務幹事(事務局), Society
  • Jan. 2026 - Present
    日本動物分類学会, 評議員, Society
  • Nov. 2024 - Present
    日本動物学会, 北海道支部 支部委員, Society
  • Jun. 2024 - Present
    日本動物学会, 第97回大会実行委員
  • May 2023 - Present
    World Register of Marine Species (WoRMS), Taxonomic editor (Tanaidacea), Others
  • Apr. 2024 - Dec. 2025
    The Japanese Society of Systematic Zoology, 選挙管理委員会, Society
  • Jul. 2017 - Nov. 2022
    日本動物学会, 学会将来計画委員, Society
  • Jul. 2018 - Sep. 2022
    日本動物学会, 北海道支部 支部委員, Society
  • May 2016 - May 2021
    日本ベントス学会, 和文誌編集委員, Society
  • Jan. 2016 - Dec. 2019
    日本動物分類学会, 英文誌編集幹事, Society
  • Jan. 2016 - Dec. 2019
    日本動物分類学会, 評議員, Society
  • Aug. 2017 - Dec. 2018
    日本動物学会, 第89回大会実行委員, Society
  • Apr. 2016 - Mar. 2018
    日本ベントス学会, 自然環境保全委員会 会計監査, Society
  • Jun. 2016 - Jun. 2016
    日本動物分類学会, 第52回大会実行委員長, Society

Research activity information

■ Awards
  • Oct. 2012, 日本動物学会, 平成24年度川口賞(川口四郎基金による若手研究者国際会議渡航補助)
    角井敬知
■ Papers
  • Additional remarks on Deltanthura palpus (Isopoda: Anthuroidea: Paranthuridae), with an amended diagnosis for Deltanthura and a note on the ontogeny of pereopod 7
    SHOKI SHIRAKI; MICHITAKA SHIMOMURA; TAEKO KIMURA; KEIICHI KAKUI
    Zootaxa, 5793, 1, 267, 278, Magnolia Press, 15 Apr. 2026, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal, Deltanthura Shiraki, Shimomura & Kakui, 2022 is a monotypic paranthurid genus established for Deltanthura palpus Shiraki, Shimomura & Kakui, 2022, which was described from a single female lacking oostegites. It was concluded from the original description that D. palpus is neotenous, because the specimen lacked pereopod 7. Recently collected material revealed that the specimen originally described was a manca-stage individual, and that the condition of pereopod 7 differed among the ontogenetic stages: it is absent in smaller specimens (6.84–10.92 mm; mancae); it has only a few setae and a tiny unguis in middle-sized specimens (12.87–13.67 mm; smaller female-like specimens and a subadult male); and it is generally similar in shape to a pereopod 6 with many setae and a normal-sized unguis, but shorter and narrower (less than 0.7 times pereopod-6 length) in larger specimens (16.56–21.14 mm; a larger female-like specimen and adult males). We present an amended diagnosis for the genus and the amended key to the genera in Paranthuridae, suggest that primary males may be present in D. palpus, and briefly discuss the possibility that this species may be undergoing an evolutionary process toward the loss of pereopod 7 (true neoteny).
  • On the capacity for desiccation-tolerant egg production in the ostracod Heterocypris spadix
    Keiichi Kakui; Mizuho Munakata
    Aquatic Animals, 2026, AA2026-21, 22 Mar. 2026, [Peer-reviewed], [Lead author, Corresponding author], [Domestic magazines]
    Japanese, Scientific journal
  • A new Nectonema nematomorph species parasitic in an alpheid shrimp (Decapoda: Caridea)
    Keiichi Kakui; Shoki Shiraki
    Journal of Helminthology, 100, e23, Cambridge University Press (CUP), 27 Feb. 2026, [Peer-reviewed], [Lead author, Corresponding author], [International Magazine]
    English, Scientific journal, Abstract

    We describe the marine horsehair worm Nectonema shimadai sp. nov. , which was parasitic in a snapping shrimp ( Alpheus sp.) collected at 4 m depth from the coastal water of Kikaijima Island, Japan, northwestern Pacific. The nematomorph was a female, 49 mm in length and 340 μm in width, salmon-pink when alive, with a translucent anterior body region and a distinct anterior chamber. The 18S sequence from N. shimadai sp. nov. was 12.8%, 9.8%, 9.6%, and 9.3% divergent in Kimura 2-parametar (K2P) distance from Nectonema sp. parasitic in the isopod Natatolana japonensis , Nectonema sp. parasitic in the crab Chionoecetes bairdi , N. munidae , and N. agile , respectively. In an 18S tree, N. shimadai sp. nov. was the sister taxon to a clade comprising three decapod-parasitic species, and species sharing the same host group or ocean were not monophyletic. The tree may reflect genetically distant warm- and cold-water lineages within decapod-parasitic nematomorphs., 51027794
  • Rhizorhina hamatipedicola n. sp. (Copepoda: Siphonostomatoida: Nicothoidae) parasitic on a tanaidacean crustacean, with a note on the effectiveness of COI and 18S data in Rhizorhina taxonomy
    Keiichi Kakui
    Systematic Parasitology, 103, 6, Springer Science and Business Media LLC, 21 Jan. 2026, [Peer-reviewed], [Lead author, Corresponding author], [Internationally co-authored], [International Magazine]
    English, Scientific journal, 51027794;11341431
  • A new species of Tropichelura (Crustacea: Amphipoda: Cheluridae) from the Ryukyu Islands, Japan
    Daigo Tokumori; Keiichi Kakui; Ko Tomikawa
    Journal of Natural History, 59, 45-48, 2657, 2672, Informa UK Limited, 08 Dec. 2025, [Peer-reviewed], [International Magazine]
    English, Scientific journal, 10695064;11341430
  • A new bopyrid species (Crustacea: Isopoda: Epicaridea: Bopyroidea) parasitic on a squat lobster from Japan, with molecular phylogenies for Epicaridea
    Yurina Katsuragawa; Shoki Shiraki; Michitaka Shimomura; Keiichi Kakui
    Marine Biodiversity, 55, 6, 118, Springer Science and Business Media LLC, 04 Dec. 2025, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal
  • A new species of the sea spider genus Cheilopallene from Japan (Pycnogonida, Callipallenidae)
    Yoshie Takahashi; Keiichi Kakui
    Spixiana, 48, 9, 13, 18 Nov. 2025, [Peer-reviewed], [Last author, Corresponding author], [International Magazine]
    English, Scientific journal
  • Description and Phylogenetic Position of Thermocypris isabella gen. et sp. nov. (Ostracoda: Cyprididae: Cyprinotinae) from a Hot Spring in Peru
    Mizuho Munakata; Hayato Tanaka; Ko Tomikawa; Yoshimi Kawasaki; Alfonso Miranda; Nilton Deza; Keiichi Kakui
    Zoological Science, 42, 5, 510, 520, Zoological Society of Japan, Oct. 2025, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal
  • Taxonomic revision of Sauranthura (Isopoda: Anthuroidea: Anthuridae), with the description of Sauranthura liangaensis sp. nov. from Surigao del Sur, Philippines
    Shoki Shiraki; Keiichi Kakui; Archie Along; Laurence Calagui; Shirlamaine Irina Calagui; Meljan Demetillo; Hiroshi Kajihara; Julius Anthony Leones; Amor Rojas-Salinas; Kris Angeli Sanchez-De los Reyes; Romell Seronay; Aoi Tsuyuki; Jamael Abato
    Marine Biology Research, 21, 4-5, 206, 220, 27 Sep. 2025, [Peer-reviewed], [Internationally co-authored], [International Magazine]
    English, Scientific journal
  • A new anthurid Amakusanthura camiguinensis sp. n. (Crustacea: Isopoda: Anthuroidea) from Camiguin Island, Philippines
    Shiraki S.; Kakui K.; Abatay M.; Caneos W.; Dahilog-Caralde M.; Kajihara H.; Tsuyuki A.; Abato J.
    Journal of Natural History, 59, 2175, 2186, 02 Sep. 2025, [Peer-reviewed], [Internationally co-authored], [International Magazine]
    English, Scientific journal
  • Intracellular Localization of the Bacterial Endosymbiont Cardinium in the Ostracod Heterocypris spadix
    Kohei Oguchi; Mizuho Munakata; Chizue Hiruta; Keiichi Kakui
    Zoological Science, 42, 4, 360, 365, Zoological Society of Japan, 31 Aug. 2025, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal
  • Overlooked Parasite Diversity in Staurozoa: Two Species of Lepocreadiidae (Trematoda: Digenea) Parasitic in Haliclystus tenuis
    Kohta Tsutsui; Keiichi Kakui
    Zoological Science, 42, 4, 352, 359, Zoological Society of Japan, 31 Aug. 2025, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal
  • Atlantapseudes tridens sp. nov. (Tanaidacea: Apseudidae) from the Okinawa Trough, Japan, with a note on the taxonomic status of Atlantapseudes curvatus
    Yoshinobu Matsushima; Maxim Jean Bâlcu; Keiichi Kakui
    Zootaxa, 5679, 484, 500, 15 Aug. 2025, [Peer-reviewed], [Last author], [Internationally co-authored], [International Magazine]
    English, Scientific journal, 13264296;11341431;10432430
  • First report of Paranthura japonica Richardson, 1909 (Crustacea: Isopoda: Paranthuridae) from Okinawa, Japan
    Shoki Shiraki; Keiichi Kakui
    Aquatic Animals, 2025, AA2025-30, 24 Jul. 2025, [Peer-reviewed], [Last author], [International Magazine]
    Japanese, Scientific journal
  • First Record of the Deep-Sea Sea Spider Bathypallenopsis californica (Arthropoda: Pycnogonida) from the Northwestern Pacific, with a Note on an Attached Crinoid
    Shogo Sekiguchi; Hisanori Kohtsuka; Keiichi Kakui
    Zoological Science, 42, 3, 335, 341, Zoological Society of Japan, 01 Jun. 2025, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal, 39595807;13264296
  • Four species of Mesanthura (Crustacea: Isopoda: Anthuroidea) from Japan, including the description of a new species
    Shoki Shiraki; Keiichi Kakui
    Journal of the Marine Biological Association of the United Kingdom, 105, e48, May 2025, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal
  • Review of the Gnathiidae (Crustacea: Isopoda) of western Japan with a description of five new species, one redescription, and one new Japanese record
    Yuzo Ota; Keiichi Kakui; Michitaka Shimomura
    Bulletin of Marine Science, 101, 2, 941, 993, Bulletin of Marine Science, May 2025, [Peer-reviewed], [International Magazine]
    English, Scientific journal, The gnathiid fauna around the world, including Japan, remains poorly detailed in the literature. In this study, we examine gnathiid specimens obtained from the continental shelf area of western Japan from 1995 to 2023. As a result, we report the following findings. (1) Two new species of Caecognathia: C. tida sp. nov., C. inferoculus sp. nov. (2) Three new species of Gnathia: G. hayashiae sp. nov., G. toyoshiomaruae sp. nov., and G. tajirana sp. nov. (3) Redescription of G. consobrina (Monod, 1926) comb. nov. since the original description. (4) First record of G. koreana Song and Ming, 2018 from Japan. Most descriptions are based on male specimens but female and third stage praniza larva are described in G. hayashiae sp. nov. New distributional records are given for G. camuripenis Tanaka, 2004, G. kumejimensis Ota, 2012 and Thaumastognathia tanseimaruae Shimomura and Tanaka, 2008. Keys to genera of Gnathiidae and the Japanese species of Caecognathia and Gnathia are provided., 10432430
  • Integrative taxonomy of Janirella (Isopoda: Asellota: Janirellidae), with descriptions of two new species from the Japan Trench and an amended classification of the genus
    Mizuki Ohta; Keiichi Kakui; Shigeaki Kojima; Michitaka Shimomura
    Biologia, 80, 6, 1407, 1423, Springer Science and Business Media LLC, May 2025, [Peer-reviewed], [International Magazine]
    English, Scientific journal, 39595807
  • Biological notes on Makassaritanais itoi (Ishimaru, 1985) (Crustacea: Peracarida: Tanaidacea)
    Keiichi Kakui
    Aquatic Animals, 2025, AA2025-11, 06 Mar. 2025, [Peer-reviewed], [Lead author, Corresponding author], [International Magazine]
    English, Scientific journal
  • Fecampiid flatworms parasitic in a tanaidacean crustacean
    Keiichi Kakui; Shoki Shiraki; Natsumi Hookabe
    Journal of Helminthology, 99, e24, 10 Feb. 2025, [Peer-reviewed], [Lead author, Corresponding author], [International Magazine]
    English, Scientific journal
  • A new genus and species of the spring-endemic Ostracoda (Cypricercinae, Cyprididae) and its genetic population structure among rheocrenic springs in Japan
    Mizuho Munakata; Hayato Tanaka; Keiichi Kakui
    Zoological Studies, 63, 51, 31 Dec. 2024, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal, We describe the ostracod Lissostrandesia fonticola gen. et sp. nov. in the subfamily Cypricercinae McKenzie, 1971, collected from six rheocrenic springs in Japan. The populations sampled were separated by up to 1000 km, and some of them by one or two marine straits, which comprise significant barriers for freshwater animals. Lissostrandesia differs from the other 13 cypricercine genera in (1) lacking a groove and inner list on the anterior inner margin of the left valve; (2) having b and d setae on the fifth limb; (3) having d1 and d2 setae on the sixth limb; (4) having a stout attachment of the caudal ramus; (5) having a Triebel’s loop on the dorsal branch of the attachment; and (6) having a long free ventral branch, its length more than twice its width, and (7) having a free dorsal branch contributing to a tip on Triebel’s loop. These differences were enough to warrant establishment of a new tribe, Lissostrandesiini, to accommodate the new genus. We present a key to the genera in Cypricercinae. The maximum p-distance for the mitochondrial cytochrome c oxidase subunit I (COI) gene among six local populations was 0.662%, a low value indicative of conspecificity. In addition, four populations with sample sizes of 13–21 individuals shared two main COI haplotypes, indicating high apparent connectivity. A trend of decreasing genetic diversity from south to north suggests L. fonticola has had a longer history on Honshu Island and dispersed northward from there. Using the 16S rRNA gene as a marker, we detected the endosymbiotic bacterium Cardinium, a group of “reproduction-manipulating” bacteria, in five populations, suggesting that L. fonticola is parthenogenetic. Passive dispersal is the most likely explanation for the broad distribution of this species across strong geographic barriers.
  • Two new Rhynchothorax sea spiders from Japan
    Hiroki Matsushita; Keiichi Kakui
    Spixiana, 47, 1, 15, 28, 17 Dec. 2024, [Peer-reviewed], [Last author, Corresponding author], [International Magazine]
    English, Scientific journal
  • Benthic deep-sea fauna in the Sea of Kumano, Japan, survey conducted from the TR/V Seisui-maru of Mie University in 2023
    Kimura T.; Kimura S.; Fujimoto S.; Kushida Y.; Tsuyuki A.; Hookabe N.; Shimooka S.; Jimi N.; Shiraki S.; Nakajima H.; Ogawa A.; Deng Z.; Kohtsuka H.; Kise H.; Kakui K.; Matsushita H.; Gregorius Altius Pratama; Kobayashi I, Hu P.; Maekawa Y.; Nakamura T.; Okumura J.; Takano M.
    The bulletin of the Graduate School of Bioresources Mie University, 50, 9, 59, Dec. 2024, [Domestic magazines]
    Japanese, Research institution
  • Colonial Entoproct Epibiotic on a Sea Spider
    Keiichi Kakui; Shogo Sekiguchi
    Zoological Science, 41, 6, 529, 532, Zoological Society of Japan, Dec. 2024, [Peer-reviewed], [Lead author, Corresponding author], [International Magazine]
    English, Scientific journal
  • Paranthura oriens sp. nov. (Isopoda: Anthuroidea: Paranthuridae) from Tateyama, Chiba, Japan
    Shoki Shiraki; Ryuta Yoshida; Keiichi Kakui
    Plankton and Benthos Research, 19, 4, 233, 243, The Plankton Society of Japan/The Japanese Association of Benthology, 28 Nov. 2024, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal
  • Lagenophryid ciliophorans on Japanese freshwater ostracods
    Mizuho Munakata; Keiichi Kakui
    Aquatic Animals, 2024, AA2024-28, 01 Nov. 2024, [Peer-reviewed], [Last author, Corresponding author], [International Magazine]
    English, Scientific journal
  • Nectonema horsehair worms (Nematomorpha) parasitic in the Tanner crab Chionoecetes bairdi, with a note on the relationship between host and parasite phylogeny
    K Kakui
    Diseases of Aquatic Organisms, 159, 153, 157, Inter-Research Science Center, 12 Sep. 2024, [Peer-reviewed], [Lead author, Corresponding author], [International Magazine]
    English, Scientific journal, Nectonema nematomorphs utilize marine crustacean hosts in their life cycle; 16 decapod and 1 isopod genera have been reported to date as host genera. This study reports the first case of Nectonema parasitic in the Tanner crab Chionoecetes bairdi, adding another known host genus. A single nematomorph juvenile was recovered from the body cavity of each of 2 ovigerous female crabs. A nucleotide sequence for the 18S rRNA gene (1854 bp) was determined from 1 Nectonema individual. The 18S sequence showed Kimura 2-parameter (K2P) distances of 10.0, 2.0, and 1.7% from 18S sequences from Nectonema sp. from an isopod host, N. agile, and N. munidae, respectively. In an 18S-based tree, the unknown species was the sister taxon to a clade comprising N. agile and N. munidae, both of which also utilize decapod hosts. The phylogenetic relationships among the 3 Nectonema species parasitic in decapods were not congruent with the phylogeny of the hosts, not supporting a hypothesis of nematomorph-host co-evolution.
  • Foettingeriidae (Ciliophora: Apostomatia) molecularly detected from Tanaidacea (Crustacea: Peracarida)
    Keiichi Kakui; Yoshihisa Fujita
    Protistology, 18, 3, 240, 243, Sep. 2024, [Peer-reviewed], [Lead author, Corresponding author], [International Magazine]
    English, Scientific journal, 31062673
  • Trematode metacercariae parasitic in the estuarine crustacean Cyathura muromiensis Nunomura, 1974 (Peracarida: Isopoda: Anthuroidea)
    Shoki Shiraki; Keiichi Kakui
    Parasitology International, 104, 102973, Elsevier BV, Sep. 2024, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal
  • Protecting stable biological nomenclatural systems enables universal communication: A collective international appeal
    Pedro Jiménez-Mejías; Saúl Manzano; Vinita Gowda; Frank-Thorsten Krell; Mei-Ying Lin; Santiago Martín-Bravo; Laura Martín-Torrijos; Gonzalo Nieto Feliner; Sergei L Mosyakin; Robert F C Naczi; Carmen Acedo; Inés Álvarez; Jorge V Crisci; Modesto Luceño Garcés; John Manning; Juan Carlos Moreno Saiz; A Muthama Muasya; Ricarda Riina; Andrea Sánchez Meseguer; Daniel Sánchez-Mata; Magdi S A El-Hawagry; Mohammad Amini Rad; Patrick Blandin; Michael Schmitt; Ryota Hayashi; Sangtae Kim; Anna Ronikier; Michał Ronikier; Sergio Chozas; Andreia Anjos; Guilherme Siniciato Terra Garbino; Mikhail Rogov; Manuel B Morales; Carlos Luis Leopardi Verde; Emre Çilden; Donald Shuka; Ermelinda Gjeta; Lulëzim Shuka; Marjol Meco; Spase Shumka; Abdelkader Nabil Benghanem; Djilali Tahri; Laouer Hocine; Rachid Meddour; Saidi Boubar; Salima Benhouhou; Clara Pladevall; Francisco Maiato Pedro Gonçalves; Abel Pérez-González; Ana Valeria Carranza; Anibal Oscar Prina; Darién E Prado; Diego G Gutiérrez; Elián L Guerrero; Esteban O Lavilla; Eugenia Minghetti; Federico Agnolín; Graciela Lorna Alfosno; Juan José Cantero; Julian Faivovich; Liliana Katinas; Magdalena Laurito; Marcelo Arana; Maria Cecilia Melo; Maria Jimena Ponce; Mariela Fabbroni; Pablo Edmundo Ortiz; Pablo Horacio Demaio; Pablo M Dellapé; Pablo Teta; Roberto Kiesling; Alla Aleksanyan; George Fayvush; Brian Jones; Daniel J Bickel; Eckart Håkansson; Elena Kupriyanova; Glenn Shea; Jürgen Kellermann; Karen L Wilson; Ladislav Mucina; Nicholas Morris; Peter Cranston; Peter Crossing; Peter G Allsopp; Peter Weston; Raymond Hoser; Rohan Pethiyagoda; Scott Eipper; Trevor Henry Worthy; Vratislav Ricardo Bejsak-Colloredo-Mansfeld; Božo Frajman; Clemens Pachschwöll; Gerald M Schneeweiss; Gerhard Pils; Hermann Voglmayr; Irmgard Krisai-Greilhuber; Konstantina Agiadi; Michael H J Barfuss; Peter Schönswetter; Wolfgang Willner; Alexander Lukashuk; Valery N Tikhomirov; Achik Dorchin; Arnaud Henrard; De Blauwe Hans; Filip Verloove; Guy Smagghe; Jacques Verniers; Jan Bosselaers; Jérôme Degreef; Kenny Meganck; Lars Chatrou; Louis Beyens; Luis M da Costa; Marc Sosef; Olivier S G Pauwels; Paolo Rosa; Paul Schoolmeesters; Pol Limbourg; Robbert Gradstein; Robert Speijer; Salvator Ntore; Thomas James Wood; Willy De Prins; Nourou Soulemane Yorou; Jigme Wangchuk; Lizette Siles; Luis F Aguirre; Luis Fernando Aguirre Urioste; Magaly Mercado U; Marita A Pareses-Rodríguez; Mauricio Ocampo; Pamela Jenny Carvajal Bacarreza; Avdul Adrović; Sadbera Trožić-Borovac; Demel Teketay Fanta; Adriano Brilhante Kury; André Hoffmann; André Silva Fernandes; Axel M Katz; Bruno Machado Teles Walter; Clapton Olimpio de Moura; Cristiano Lopes-Andrade; Dalton de Souza Amorim; Daniel C Cavallari; Daniel Loebmann; Diego Aguilar Fachin; Diego Nunes da Silva; Fernando Z Vaz-de-Mello; Francisco Langeani Neto; Gerson Paulino Lopes; Henrique C Costa; Henrique Mallmann Büneker; Isabel Paiola Gonçalves; John E Lattke; Jose Albertino Rafael; José Nascimento; José Rubens Pirani; Josimar Külkamp; Julian C G Silva Junior; Katyuscia Araujo-Vieira; Leandro Lacerda Giacomin; Leandro Manzoni Vieira; Leandro Tavares Azevedo Vieira; Leonardo M Borges; Leonardo M Versieux; Lucas Cardoso Marinho; Lucas Sá Barreto Jordão; Luciano Paganucci de Queiroz; Marcelo Trovó; Marcelo Weksler; Marcos Gonçalves Lhano; Maria Beatriz Rossi Caruzo; Marinus S Hoogmoed; Maurício Figueira; Miguel Medrano; Milton Groppo; Omar Rojas Padilla; Otávio Luis Marques da Silva; Patricia Jacqueline Thyssen; Pedro Henrique Cardoso; Rafael A P Freitas Silva; Renata Trevizan; Ricardo de S Secco; Rosângela Simão-Bianchini; Scott Thomsom; Tamily Carvalho Melo dos Santos; Ulisses Caramaschi; Vinícius Sudré dos Santos; Weferson Júnio da Graça; Yuri Rossine; Johan Willem Frederik Slik; Ulmar Grafe; Anita Tosheva; Asen Asenov; Dolja Pavlova; Fedor Konstantinov; Georgi Kunev; Nikolay Simov; Alain Didier Missoup; Andrew M R Bennett; Brenda Hann; Chad Morgan; Étienne Lacroix-Carignan; Étienne Léveillé-Bourret; Felix Sperling; Fenja Brodo; Geraldine A Allen; Gregory R Pohl; James Basinger; Jean-François Landry; John E Maunder; John Huber; John Warren Reynolds; Jonathan Cazabonne; Juan Carlos Villarreal Aguilar; Matthias Buck; Mihai Costea; Olivier Barden; Quentin Cronk; Rafael Otfinowski; Santiago Claramunt; Stephen Darbyshire; Timothy Dickinson; William J Crins; Alicia Marticorena Garri; Diego Alarcón Abarca; Francisco A Squeo; Guillermo D'Elía; Jorge Pérez-Schultheiss; José Angel García-Beltrán; Marcelo M Rivadeneira; Sara M Rodríguez; Sebastian Teillier; Sven N Nielsen; Amael Borzee; Baojun Zhao; Bismillah Shah; Cai-Xia Yuan; Caixin Liu; Chen-Yan Liao; Chengyuan Liu; Chuan-Bu Gao; Chufei Tang; Chunhao Zhang; Clive Siu-Ki Lau; Dakang Zhou; Daniel Roland Gustafsson; Dávid Rédei; Fang-Meng Duan; Fenglong Jia; Fuqiang Chen; Guchun Zhou; Gui-Qiang Huang; Hao Xu; Hao-Ran Gao; Haodong Yan; Hassan Naveed; Hongbin Zhu; Hongliang Shi; Hu Li; Huilin Han; Ji-Liang Liu; Jia Huang; Jiabin Liu; Jian qiang Cao; Jian Sun; Jian-Wei Guo; Jian-Yu Chen; Jianjun Guo; Jiao Sun; Jiaxin Wang; Jing-yu Zhai; Jing-zhi Lin; Jinrui Zhang; Jun Chen; Jun Zhang; Junbo Tong; Kai Kang; Kai Wang; Kai-Ming Cai; Keping Ma; Kexin Xu; Kuiyan Zhang; Li Ren; Liang Tang; Lin Luo; Ling-Zeng Meng; Mei-Ying Lin; Meng Xie; Mingyu Liu; Muhammad Asghar Hassan; Panshuai Fei; Patrick Rioual; Peng Li; Pipeng Li; Prabin Bhandari; Qingshuo Xu; Qionghua Gao; QiRong GUO; Rongrong Shen; Runcheng Huang; Sheng Gao; Shi-Yang Feng; Shu Li; Shuang Qiu; Shuang Wang; Shujie Zhang; Shuren Zhang; Siqin Ge; Siyuan Xu; Tao Yang; Tian-Hao Hu; Tian-Long He; Tiangang Gao; Wenqi Yin; Xiang-qian Sun; Xiao-Feng Jin; Xiao-Ran Yang; Xiaoqi Zeng; Xin-Lei Zhao; Xin-Ran Li; Xing-Ke Yang; Xing-Long Bai; Xingyue Liu; Xinle Duan; Xinyi Zheng; Xiumin Li; Xu Zhang; Xue Jiang; Yahui Zhao; Yanyan Zhao; Yi-Chen Fan; Yi-Fei Lu; Yi-Feng Zhang; Yifeng Liu; Yiming Yu; Yingxin Zhang; Yong Zhou; Yongke Zhang; Yongsheng Yan; You-Wen Zhao; Yu-jian Li; Yuting Dai; Zehai Hou; Zhao Pan; 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Alexey S Shmakov; Alexey V Lopatin; Alisa A Neplyukhina; Andrei A Legalov; Andrey Samdan; Andrey V Matalin; Andrey V Zhuravlev; Artem Leostrin; Artem Y Sinev; Boris A Korotyaev; Boris Kataev; Davidian Genrik; Dmitry A Lifanov; Dmitry Fedorenko; Dmitry Kopylov; Dmitry Lyskov; Dmitry Sokoloff; Dmitry Vorontsov; Eduard V Mychko; Efremov Andrey; Egorov Leonid; Evgeniy Dunayev; Evgeniya Muchnik; Evgeny V Popov; Gennady Baryshnikov; Georgy V Mirantsev; Gyulli Farzalieva; Igor Fefelov; Irina Samsonova; Julia Shuvalova (Yulia/Yuliya Shuvalova); Kirill Mikhailov; Kirill V Makarov; Konstantin K Tarasenko; Ksenia G Klimova; Larisa Bagmet; Lev Shestakov; Lovtsova Julia; Lyudmila Mikhailovna Pavlova; Maxim Nabozhenko; Maxim Nuraliev; Maxim Vinarski; Melnikov Denis G; Michael Serebryanyi; Natalia Terekhina; Natalya Andreyenkova; Nikita Zdravchev; Nikita Zelenkov; Nikolay A Poyarkov; Pavel Khramtsov; Pavel Parkhaev; Peter Khramov; Roman Nazarov; Sergei E Tshernyshev; Sergey Sinev; Vladimir Fedosov; Vladislav Desiatkin; Yuri Mikhailov; Yurii N Sundukov; Zinovyev Evgeniy; Ahmed M Soliman; Abdoulaye Baïla Ndiaye; Mingou Paterne Arnaud Bernard; Aleksandar Ćetković; Anđeljko Petrović; Dalibor Z Stojanović; Mihailo Vujić; Mirko Šević; Predrag Simonović; Saša Marić; Vera Nikolic; Vukašin Gojšina; John S Ascher; Adam Kantor; Anna Guttova; Daniel Jablonski; Judita Zozomová; Karol Marhold; Pavol Mereďa; Stanislav Španiel; Vladimír Hemala; Andraž Čarni; Faisal Jama Gelle; Benny Bytebier; Christian M Deschodt; David Gwynne-Evans; Denis Brothers; Gimo M Daniel; Hester Steyn; Ian Engelbrecht; Jacques van Rooy; Janine Victor; Janneke Aylward; Lynne J Quick; Mike Mostovski; Musa C Mlambo; Riaan Stals; Ruan van Mazijk; Susan Dippenaar; Vinícius da Costa-Silva; A Jesús Muñoz-Pajares; Aaron Pérez-Haase; Agustin Lahora; Alberto Rodríguez-Fernández; Ana Beleén Fernández Salegui; Ana García-Muñoz; Ana María Vega Maray; Ana Morales-Alonso; Ana Ortega-Olivencia; Andres Baselga; Andres Ordiz; Ángel Fernández Ceular; Antonio Flores-Moya; 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Llorenç Saez; Lluis Serra Laliga; Lourdes López Merino; M Luisa Buide del Real; M Mar Trigo; M Montserrat Martínez-Ortega; Magdalena Vicens Fornés; Manuel A Rodriguez-Iglesias; Manuel de la Estrella; Manuel J Macía; Manuel Sánchez-Angulo; Marcial Escudero; María Luisa Navarro Pérez; María Sanz Arnal; Mario Fernández-Mazuecos; Marta Alcolea; Miguel Ángel Gómez-Serrano; Miquel Capó; Mohamed Abdelaziz; Mónica Miguez Ríos; Nereida M Rancel-Rodríguez; Oscar Reverter-Gil; Pablo García Murillo; Pablo Muñoz-Rodríguez; Pablo Tejero Ibarra; Pablo Vargas; Paloma Cantó; Pascual López-López; Pedro Pérez Olea; Pedro Sánchez Gómez; Penélope González-Sampériz; Pere Fraga i Arguimbau; Rafael Medina; Rafael Molina Venegas; Raquel Alonso Redondo; Raquel Pino Bodas; Raúl Lois Madera; Raúl Ochoa-Hueso; Raúl Orihuela-Rivero; Raúl Sánchez Francés; Regina Berjano; Ricardo Garilleti; Rodrigo R Granjel; Rogelio Sánchez Villegas; Rosa Mª Valencia Barrera; Rosario G Gavilan; Rosina Magaña Ugarte; Rubén G Mateo; Ruymán David Cedrés-Perdomo; Santiago Andrés Sánchez; Santiago Castroviejo-Fisher; Santiago Fernández Jiménez; Santiago Ortiz; Sara del Río; Sara Romero; Sergio Pérez-Ortega; Silvia Gómez Valle; Sonia Molino; Teresa Garnatje; Tomás Rodríguez Riaño; Víctor J Rico; Victoria Ferrero Vaquero; Violeta Atienza; Violeta Simón-Porcar; Virginia Valcárcel; William Douglas Carvalho; Wolfgang Denzer; Yoana García Mendizabal; Bengt Å Bengtsson; Bengt Carlsson; Brita Svensson; Carlos Guerrero Bosagna; Eric Wahlsteen; Göran Arnqvist; Irene Bisang; Jadranka Rota; Johan Liljeblad; Kennet Lundin; Knud Andreas Jønsson; Lars Hedenäs; Mats Wedin; Michael Norén; Niklas Wahlberg; Paco Cárdenas; Per Ahlberg; Per Alström; Per Löfgren; Peter Nielsen; Sabine Stöhr; Steffen Kiel; Torbjörn Tyler; Ulf Arup; Zongzhuang Liu; Arne Strid; German Hernández Alonso; Alain Chautems; Ana B García-Martín; André Charpin; Andreas Müller; Anne Freitag; Bernard Landry; Christophe Praz; Danae Thivaiou; Daniel Cherix; Daniel Jeanmonod; Edward Mitchell; Florian Mombrial; Giulio Cuccodoro; Guido Sabatinelli; Ivan Löbl; Jean Mariaux; Jean-Paul Haenni; Jean-Paul Theurillat; Juan Carlos Zamora; Laurent Valtotton; Manuel Ruedi; Marvco V Bernasconi; Nicolas Fumeaux; Oliver Hawlitschek; Peter Schuchert; Philippe Clerc; Seraina Klopfstein; Adun Samathi; Kevin Hyde; Raoufou Radji; J Francisco Morales; Faouzia Charfi; Ridha El Mokni; Filiz Gunay; Saida Agayeva; Alessandro Giusti; André Schuiteman; Andrew E Whittington; Archie YF Ko; Ashley Leftwich; Christopher David Williams; Clare Scanlan; Crispin Little; Danniella Sherwood; David C Lees; David Goyder; David Michael John; Edward Dickinson; George W Beccaloni; Henry J Noltie; Ian F G McLean; Isabel Larridon; John Badmin; John M McArthur; John S Noyes; Jonathan A Todd; Leonidas Romanos Davranoglou; Mark R Shaw; Mark T Young; Martin Ebejer; Martyn Kelly; Maurizio Dioli; Maxwell V L Barclay; Michael F Geiser; Nick Stewart; Peter Langton; Richard M Bateman; Rob Marchant; Simon Schneider; Tabitha R Taberer; Tammy Horton; Vladimir Blagoderov; Wolfgang Wüster; Zhe-Yu Chen; Dmitry Telnov; Andriy Novikov; Dmytro Dubyna; Igor Balashov; Ihor Olshanskyi; Mykola Fedoronchuk; Natalia M Shiyan; Oleksandr Khodosovtsev; Oleksandr Kovalchuk; Oxana Vinogradova; Tatiana Mikhailyuk; Vasily Dyadichko; Vera Hayova; Viktor O Nachychko; Vitalii Kolomiichuk; Vitaliy A Kharchenko; Evgeny E Perkovsky; Ary Mailhos; Eduardo Marchesi; Jose Mauricio Bonifacino; María Victoria Valtierra Sierra; Aaron M Bauer; Adeyinka Adeyemi; Alan Franck; Alessandra Pandolfi; Allen L Norrbom; Andre E Moncrieff; Andrew Johnson; Angelo Capparella; Anthony Rossi; Armin Namayandeh; Art Borkent; Barbara Hellenthal; Brett C Ratcliffe; Brian J O'Shea; Brian Tilston Smith; C Jonathan Schmitt; Camm C Swift; Chris M Cohen; Christopher J Borkent; D Christopher Rogers; Dale Casamatta; Daniel Koenemann; David Boufford; David Campbell; David W Krause; Davide Dal Pos; Derek Sikes; Don Les; Edward S DeKeyser; Edwin Cruz-Rivera; Emily Carter; Glenn Fairbanks Seeholzer; Guy Eroh; H Douglas Pratt; Hinrich Kaiser; Irene Villa Machio; J Mark Egger; J V Remsen; Jacob Esselstyn; James Christopher Sagebiel; James Liebherr; James N Hoghe; Jared Wolfe; Jason J Dombroskie; Jessica Eberhard; Jimmy A McGuire; John A Gerwin; John H Epler; José H Leal; Jose L Lopez-Ribot; Judith E Winston; Julian Harber; Jun Wen; Justin L Lee; Justin Pargeter; Katharine Gregg; Keith Arnold; Kenneth Carpenter; Kevin McCracken; Kevin Williams; Kevin Winker; Kipling Will; Kyhl A Austin; Kyle E Harms; Les Watling; Linda Watson; Manuel Marin; Marc Edward Epstein; Marcus M Key; Mariana P Marques; Mark Pyron; Martin Dovciak; Martin Hauser; Mary Jane West-Eberhard; Mauricio Diazgranados; Mel Warren; Michael K Oliver; Michael L Ferro; Michael S Engel; Natalia von Ellenrieder; Nathan Burkett-Cadena; Neal L Evenhuis; Neil Snow; Nobuaki Mizumoto; Obediah Sage; Patrick Rakes; Patrick Rohner; Paul J Morris; Paula Cushing; Paula Rozo-Lopez; Paulo Durães Pereira Pinheiro; Peter H Kerr; Peter Stevens; Peter Uetz; Quentin Wheeler; R Edward DeWalt; R Kevan Schoonover McClelland; Richard H Zander; Richard Rabeler; Robert L Zuparko; Robert W Hansen; Roberta Brett; Ronald Hellenthal; Rose Meier; Rosser W Garrison; Roy E Gereau; Sam W Heads; Samuel A McLeod; Samuel Wooster James; Severyn Korneyev; Shaun L Winterton; Stephen D Gaimari; Steve Hilty; Subir Shakya; Susan B Adams; Susan G Letcher; Thomas H Atkinson; Timothy S Wood; Victor H Gonzalez; Walsh SJ; Wendy Applequist; William F Smith-Vaniz; William L Murphy; William Wayt Thomas; Khalimov F Z; Mukhammadtuychi Rakhimov; Shodmonov Feruz Abdullajonovich; Yigitali Tashpulatov; Fernando J M Rojas-Runjaic; Hugo Gullermo Cerda Pérez; Pedro Torrecilla; Thirza Ruiz Zapata; Hoang Quynh Hoa; Ngoc Nguyen; Cathy Sharp; Moira Fitzpatrick
    BioScience, 74, 7, 467, 472, Oxford University Press (OUP), 16 Aug. 2024, [Peer-reviewed], [Internationally co-authored], [International Magazine]
    English, Scientific journal, Abstract

    The fundamental value of universal nomenclatural systems in biology is that they enable unambiguous scientific communication. However, the stability of these systems is threatened by recent discussions asking for a fairer nomenclature, raising the possibility of bulk revision processes for “inappropriate” names. It is evident that such proposals come from very deep feelings, but we show how they can irreparably damage the foundation of biological communication and, in turn, the sciences that depend on it. There are four essential consequences of objective codes of nomenclature: universality, stability, neutrality, and transculturality. These codes provide fair and impartial guides to the principles governing biological nomenclature and allow unambiguous universal communication in biology. Accordingly, no subjective proposals should be allowed to undermine them.
  • Isopods on isopods: integrative taxonomy of Cabiropidae (Isopoda: Epicaridea: Cryptoniscoidea) parasitic on anthuroid isopods, with descriptions of a new genus and three new species from Japan
    Shoki Shiraki; Keiichi Kakui
    Invertebrate Systematics, 38, IS24013, 01 Aug. 2024, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal
  • Apseudes ranma sp. nov. (Tanaidacea: Apseudidae) found in a public aquarium, with notes on phylogeny and a presumptive stridulatory organ
    Yoshinobu Matsushima; Keiichi Kakui
    Bulletin of Marine Science, 100, 451, 469, Bulletin of Marine Science, 25 Jul. 2024, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal, We describe Apseudes ranma sp. nov. from the Port of Nagoya Public Aquarium, Nagoya, Japan. It resembles two congeners, Apseudes curtiramus (Guţu, 2007) and Apseudes orghidani Guţu and Iliffe, 1989, in having the rostrum triangular, wider than long, the carapace without lateral spines, distinct and round eye lobes, pereonites 3–6 with a pair of tongue-shaped anterolateral projections, and the pleotelson with two pairs of lateral projections, but differs from them in having lateral setae on the anterolateral margin of the carapace and a ventro-subproximal spine on the pereopod-1 basis, and in the number of spiniform setae on articles of pereopods 2–4. We determined partial nucleotide sequences for the cytochrome c oxidase subunit I (COI) and 18S rRNA (18S) genes in A. ranma sp. nov. for future DNA barcoding and phylogeny reconstruction. In an 18S tree, A. ranma sp. nov. was the sister taxon to Apseudes nipponicus Shiino, 1937, and this clade was the sister group to moderately supported Carpoapseudes clade. We discovered a presumptive stridulatory organ in A. ranma sp. nov. consisting of a cylindrical protrusion and knobbly area on the inner surface of each of the left and right chelipedal bases. The location of the protrusion and knobbly area suggests that these may be a plectrum and pars stridens, respectively., 13264296;11341431;10432430
  • Ontogenetic Changes in Pigmentation Pattern in Mesanthura Miyakoensis (Crustacea: Isopoda: Anthuridae)
    Shoki Shiraki; Keiichi Kakui
    Zoological Science, 41, 3, 323, 328, Zoological Society of Japan, 01 Jun. 2024, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal
  • Brixitanais, a new substitution name for Brixia Jóźwiak, Drumm, Bird & Błażewicz, 2018 (Crustacea: Tanaidacea: Akanthophoreidae)
    Keiichi Kakui
    Zootaxa, 5458, 3, 449, 450, Magnolia Press, 29 May 2024, [Peer-reviewed], [Lead author, Corresponding author], [International Magazine]
    English, Scientific journal
  • Description of a new Cruranthura species from the Miyako Islands, Japan, with its DNA barcode (Isopoda: Anthuroidea: Paranthuridae)
    Shoki Shiraki; Keiichi Kakui
    Species Diversity, 29, 1, 65, 72, Japanese Society of Systematic Zoology, 25 May 2024, [Peer-reviewed], [Last author], [International Magazine]
    English, Scientific journal
  • A new parasitic barnacle (Crustacea, Cirripedia, Rhizocephala, Mycetomorpha) from the abyssal zone in the northwestern Pacific
    Keiichi Kakui
    Zoosystematics and Evolution, 100, 2, 385, 390, Pensoft Publishers, 03 Apr. 2024, [Peer-reviewed], [Lead author, Corresponding author], [International Magazine]
    English, Scientific journal, I describe the parasitic barnacle Mycetomorpha abyssalissp. nov. from the crangonid shrimp Sclerocrangon zenkevitchi collected from 3893–3890 m depth off the eastern coast of Iwate, Japan, northwestern Pacific. This is the first Mycetomorpha rhizocephalan from the abyssal zone and the third species in Mycetomorpha. Mycetomorpha abyssalissp. nov. differs from its congeners M. vancouverensis and M. albatrossi in (1) triangular shield lacking, (2) stalk one-quarter of length from posterior end of externa, (3) mantle opening clearly anterior to stalk, (4) different host genus, and (5) depth range much deeper. I determined partial sequences for the mitochondrial cytochrome c oxidase subunit I (COI) and 16S rRNA genes and nuclear 18S rRNA and 28S rRNA genes from M. abyssalissp. nov. for future DNA barcoding and phylogeny. Kimura 2-parameter distances between M. abyssalissp. nov. and M. vancouverensis were 21.2% (16S), 0.6% (18S), and 1.5% (28S)., 39595807;13264296
  • Molecular phylogeny confirms that “Pseudolepeophtheirus” schmidti Gusev, 1951 belongs in Lepeophtheirus
    Keiichi Kakui
    Spixiana, 46, 2, 173, 177, Apr. 2024, [Peer-reviewed], [Lead author, Corresponding author], [International Magazine]
    English, Scientific journal
  • Review on studies of fossils in Tanaidacea (Crustacea: Malacostraca: Peracarida)
    Keiichi Kakui
    Fossils, 115, 65, 76, 31 Mar. 2024, [Peer-reviewed], [Lead author, Corresponding author], [Domestic magazines]
    Japanese, Scientific journal
  • Marine macroinvertebrate fauna of the shallow coastal waters of the Shiretoko Peninsula, a World Natural Heritage at Hokkaido, Japan
    Takeshi Sonoda; Susumu Chiba; Tomoyasu Yamazaki; Takahiro Nobetsu; Hideaki Fujiya; Tomoyuki Komai; Noboru Nunomura; Michitaka Shimomura; Ko Tomikawa; Keiichi Kakui; Takashi Hosono; Atsushi Ito; Toshihiko Fujita; Itaru Kobayashi; Seiji Goshima
    Plankton and Benthos Research, 19, 1, 1, 16, The Plankton Society of Japan/The Japanese Association of Benthology, 28 Feb. 2024, [Peer-reviewed], [International Magazine]
    English, Scientific journal
  • A smaller species releases proportionally larger juveniles in Apseudes (Crustacea: Peracarida: Tanaidacea)
    Keiichi Kakui; Chizue Hiruta
    CBM-Cahiers de Biologie Marine, 65, 35, 37, Station Biologique de Roscoff (SBR), 26 Jan. 2024, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, We compared the body size (carapace width, CW) of the first free-living instar individuals, or mancae 2, and of the smallest ovigerous individuals, between two simultaneously hermaphroditic tanaidacean species, Apseudes nipponicus Shiino, 1937 and Apseudes sp. The former species is 2.2 times larger, with a CW of 2.61 mm in the smallest ovigerous individual, compared to 1.17 mm in the latter. Average CWs of manca 2 individuals were 0.51 mm (A. nipponicus, n = 3, 0.50-0.51 mm) and 0.38 mm (Apseudes sp.: n = 9, 0.38-0.39 mm), meaning that A. nipponicus mancae 2 were 20% as large as the smallest ovigerous individuals, whereas those of Apseudes sp. were 32% as large. The proportionally larger juveniles in the smaller species are consistent with the progenetic hypothesis that Apseudes sp. is achieving earlier maturation as females., 13264296
  • Flatworm cocoons in the abyss: same plan under pressure
    Keiichi Kakui; Aoi Tsuyuki
    Biology Letters, 20, 20230506, 24 Jan. 2024, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 39595807;13264296
  • A New Species of Coralliidae (Cnidaria: Octocorallia) Collected from Eastern Japan
    Masanori Nonaka; Nozomi Hanahara; Keiichi Kakui
    Species Diversity, 28, 2, 231, 243, Japanese Society of Systematic Zoology, 09 Nov. 2023, [Peer-reviewed], [Last author]
    English, Scientific journal
  • A new species of free-living marine nematode, Fotolaimus cavus sp. nov. (Nematoda, Oncholaimida, Oncholaimidae), isolated from a submarine anchialine cave in the Ryukyu Islands, southwestern Japan
    Shimada D; Kakui K; Fujita Y
    Zoosystematics and Evolution, 99, 2, 519, 533, Pensoft Publishers, 06 Nov. 2023, [Peer-reviewed]
    Japanese, Scientific journal, Fotolaimus cavussp. nov. was described from a submarine anchialine cave called Akuma-no-yakata on the Shimoji Island, Miyako Island Group, Ryukyu Islands, southwestern Japan. This is the first free-living marine nematode isolated from a submarine cave in Japan, and the third species of the genus Fotolaimus. This new species differs from its congeners by its small body size, wide amphids, long buccal cavity, long conico-cylindrical tail, and proximally curved gubernaculum. We provide amended dichotomous keys to genera in the subfamily Oncholaiminae and species in Fotolaimus. We also analyzed partial DNA sequences encoding ribosomal small subunit RNA and cytochrome c oxidase subunit I from Fotolaimus cavussp. nov. and six other species of Oncholaimidae collected from Japanese waters. The phylogenetic tree based on the ribosomal small subunit RNA sequences using maximum likelihood analysis suggested a close relationship between Fotolaimus and Wiesoncholaimus as well as Oncholaimus. The topology of the tree was similar to those from previous studies; however, it suggested a new phylogenetic position of Adoncholaimus as a sister clade for Viscosia and Oncholaimus., 31062673
  • Two new species in Leptocheliidae (Crustacea: Peracarida: Tanaidacea) from Japan, with notes on their phylogenetic position and aspects of morphology
    Akari Sato; Kazuharu Arakawa; Keiichi Kakui
    Zoologischer Anzeiger, 306, 51, 68, Elsevier BV, 31 Aug. 2023, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal, 13264296;11341431
  • Capropodocerus, a new genus in Podoceridae (Crustacea: Amphipoda) from Japan, with descriptions of two new species.
    Yu Matsumoto; Hiroshi Kajihara; Keiichi Kakui
    Zootaxa, 5336, 4, 577, 589, Magnolia Press, 24 Aug. 2023, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal, We establish the new podocerid genus Capropodocerus gen. nov. for the two new species Capropodocerus tagamaru gen. et sp. nov. (collected from 775–800 m depth in the Kumano Sea, Japan) and Capropodocerus kamaitachi gen. et sp. nov. (collected from 840–873 m depth off the eastern coast of Miyagi, Japan). Capropodocerus shares elongate pereonites 1–3 with Neoxenodice but differs from the latter in lacking uropod 2 and in having pereonite 3 at least twice as long as pereonite 4. Capropodocerus is also similar to Laetmatophilus and Leipsuropus in having only two pairs of uropods but differs from them in having i) pereonite 1 longer than the head and ii) pereonite 3 at least twice as long as pereonite 4; Capropodocerus has three urosomes and Laetmatophilus has two urosomes. The two species in Capropodocerus differ in i) the length ratio of the antenna 1 peduncle to the flagellum, ii) the shape of the gnathopod 2 palm, iii) the presence or absence of a mid-posterior triangular projection on the gnathopod 2 dactylus, and iv) the length ratio of the basis to the merus of pereopod 3. A key to all genera in Podoceridae is provided., 10432431;11341431
  • Differential masculinization of the chelipeds in two simultaneously hermaphroditic Apseudes tanaidaceans (Crustacea: Malacostraca)
    Keiichi Kakui; Chizue Hiruta
    Zoologischer Anzeiger, 305, 23, 27, Elsevier BV, Jul. 2023, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 13264296
  • New sea spider species (Pycnogonida: Austrodecidae) from a submarine cave in Japan
    Keiichi Kakui; Yoshihisa Fujita
    Journal of the Marine Biological Association of the United Kingdom, 103, e44, 23 Jun. 2023, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 31062673
  • Diexanthema hakuhomaruae sp. nov. (Copepoda: Siphonostomatoida: Nicothoidae) from the Hadal Zone in the Northwestern Pacific, with an 18S Molecular Phylogeny
    Keiichi Kakui; Jun Fukuchi; Mizuki Ohta
    Acta Parasitologica, 68, 413, 419, Springer Science and Business Media LLC, 21 Jun. 2023, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 39595807;13264296
  • Phototaxis in Two Shallow-Water Zeuxo Species (Crustacea: Tanaidacea)
    Nobuya Okamoto; Keiichi Kakui
    Zoological Science, 40, 3, 203, 207, Zoological Society of Japan, 01 Jun. 2023, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal, 11341431
  • A new Sphaeronella species (Copepoda: Siphonostomatoida: Nicothoidae) parasitic on Euphilomedes sp. (Ostracoda: Myodocopa: Philomedidae) from Hokkaido, Japan, with an 18S molecular phylogeny
    Keiichi Kakui; Mizuho Munakata
    Systematic Parasitology, 100, 2, 121, 131, Springer Science and Business Media LLC, Mar. 2023, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Freshwater Ostracoda (Crustacea) from Rishiri Island, Hokkaido, Japan
    Mizuho Munakata; Shinri Tomioka; Keiichi Kakui
    Rishiri Studies, 42, 5, 18, Mar. 2023, [Last author]
    Japanese, Research institution
  • A new species of Leipsuropus Stebbing, 1899 (Amphipoda: Podoceridae) from Japan
    Yu Matsumoto; Hiroshi Kajihara; Keiichi Kakui
    Nauplius, 31, e2023005, FapUNIFESP (SciELO), 20 Feb. 2023, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal, 11341431
  • On the original publications for six tanaidacean names proposed by Sueo M. Shiino
    MASAATSU TANAKA; KEIICHI KAKUI
    Zootaxa, 5231, 2, 192, 194, Magnolia Press, 27 Jan. 2023, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal
  • Integrative taxonomy of Zeuxo (Crustacea: Peracarida: Tanaidacea) from Japan, with the description of a new species
    Nobuya Okamoto; Keiichi Kakui
    Biologia, 77, 9, 2497, 2506, Springer Science and Business Media LLC, 31 Oct. 2022, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal, 13264296;11341431
  • Protogynous hermaphroditism in Crustacea: a new example from Tanaidacea
    Keiichi Kakui; Chizue Hiruta
    Canadian Journal of Zoology, 100, 8, 481, 487, Canadian Science Publishing, Aug. 2022, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, Hermaphroditism has been widely reported in the Crustacea, but protogyny, in which females change to males, is apparently rare and restricted to the peracarid orders Isopoda and Tanaidacea. In the latter, protogyny has been demonstrated in only seven species representing six genera by rearing experiments and indicated for several other genera and species through morphology. Here, we show through rearing experiment and histological observations, protogynous hermaphroditism in the tanaidacean species Nesotanais sp. aff. ryukyuensis Kakui, Kajihara and Mawatari, 2010. Our rearing experiment and size distribution data for wild individuals showed that females can change sex at various sizes (= ages). We found one individual identified externally as a female containing both ovaries and testes, indicating that overt female individuals undergo a short transitional phase as simultaneous hermaphrodites before the sex-change molt. We discuss the relationship among the wide size distribution of sex-changing females, the short life span of males, and the tube-dwelling mode of life., 13264296;11341431;10905851
  • First report of Hyssuridae (Isopoda: Anthuroidea) from Japan, with the description of a new Kupellonura species
    Shoki Shiraki; Michitaka Shimomura; Keiichi Kakui
    Nauplius, 30, e2022023, FapUNIFESP (SciELO), Aug. 2022, [Peer-reviewed], [Last author]
    English, Scientific journal
  • A new brackish tanaidacean, Sinelobus kisui sp. nov. (Crustacea, Peracarida, Tanaidacea), from Japan, with a key to Sinelobus species and barcode information from two loci
    Kyoko Hirano; Keiichi Kakui
    Zoosystematics and Evolution, 98, 245, 256, 29 Jun. 2022, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal, 13264296
  • Biological notes on Zeuxo ezoensis Okamoto et al., 2020 (Crustacea: Peracarida: Tanaidacea).
    Yoshinobu Hayakawa; Keiichi Kakui
    Aquatic Animals, 2022, AA2022-10, 09 Jun. 2022, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal
  • Observations on Predation in Paranthura japonica Richardson, 1909 (Isopoda: Cymothoida: Paranthuridae)
    Shoki Shiraki; Keiichi Kakui
    Zoological Science, 39, 3, 270, 274, Zoological Society of Japan, 06 Jun. 2022, [Peer-reviewed], [Last author]
    English, Scientific journal, 13264296
  • Dive into the sea: first molecular phylogenetic evidence of host expansion from terrestrial/freshwater to marine organisms in Mermithidae (Nematoda: Mermithida)
    K. Kakui; D. Shimada
    Journal of Helminthology, 96, Cambridge University Press (CUP), May 2022, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, Abstract

    We report the first mermithid nematode found to be parasitic in a marine tanaidacean crustacean. Ten host tanaidaceans were collected from a depth of 52 m in Otsuchi Bay, Iwate, Japan, north-western Pacific, and identified as a species in the tanaidid genus Zeuxo Templeton, 1840. Nematodes occurred in the host's body cavity; in one case, at least two individuals inhabited a single host. We provide a brief description and illustrations of the morphology of the nematode. In a phylogenetic reconstruction based on the 18S ribosomal RNA gene, the nematode nested in a clade otherwise containing mermithids from terrestrial or freshwater hosts, showing an expansion in host utilization in Mermithidae Braun, 1883 from terrestrial/freshwater hosts to a marine organism.
  • Taxonomy and natural history of Cavernocypris hokkaiensis sp. nov., the first ostracod reported from alpine streams in Japan
    Mizuho Munakata; Hayato Tanaka; Keiichi Kakui
    Zoosystematics and Evolution, 98, 1, 117, 127, Pensoft Publishers, 05 Apr. 2022, [Peer-reviewed], [Last author]
    English, Scientific journal, We describe the cypridoidean ostracod Cavernocypris hokkaiensis sp. nov. from riverbed sediments in an alpine stream at an elevation of ca. 1850 m in the Taisetsu Mountains, Hokkaido, Japan. This species differs from congeners in having (1) the outer surface of the carapace smooth, with sparse, tiny setae, but without shallow pits; (2) the carapace elongate rather than triangular in lateral view; (3) the antennula consisting of seven podomeres; (4) first palpal podomere of maxillula with five dorsodistal and one ventro-subdistal setae; (5) the fifth limb lacking setae b and d; and (6) the fifth limb lacking a vibratory plate. We provided the key to the Cavernocypris species. We determined partial sequences for the cytochrome c oxidase subunit I (COI; cox1) and 18S rRNA (18S) genes in C. hokkaiensis. Our sample contained only females, and we obtained a partial 16S rRNA sequence for the endosymbiotic bacterium Cardinium from C. hokkaiensis, indicating the possibility that this ostracod species reproduces parthenogenetically. Our field survey and observations of captive individuals suggested that C. hokkaiensis may be endemic to the Taisetsu Mountains, with a low population density, narrow distributional range, and slow maturation to sexual maturity.
  • Digenean Metacercariae Parasitic in a Staurozoan Cnidarian
    Keiichi Kakui
    Zoological Science, 39, 2, Zoological Society of Japan, 01 Apr. 2022, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • A new neotenous genus and species, Deltanthura palpus gen. et sp. nov. (Isopoda, Anthuroidea, Paranthuridae) from Japan, with a revised key to the genera in Paranthuridae
    Shoki Shiraki; Michitaka Shimomura; Keiichi Kakui
    Zoosystematics and Evolution, 98, 1, 109, 115, Pensoft Publishers, 31 Mar. 2022, [Peer-reviewed], [Last author]
    English, Scientific journal, We describe a new paranthurid isopod genus and species, Deltanthura palpusgen. et sp. nov., collected from a depth of 805–852 m off the southern coast of Mie prefecture, Japan. Deltanthura is similar to Pseudanthura Richardson, 1911 in having a triangular pleotelson, acute mandible with a 3-articulate palp, a maxillipedal endite, and a tapering uropodal exopod, but differs in having eyes and neotenous characters (reduced pereonite 7 and pereopods 7 lacking). Deltanthura and four paranthurid genera (Califanthura Schultz, 1977, Colanthura Richardson, 1902, Cruranthura Thomson, 1946, and Cruregens Chilton, 1882) share neotenous characters, but in Deltanthura the mandible is acute, with a 3-articulate palp and maxillipedal endites are present. Califanthura minuta Kensley & Heard, 1991 may belong in Deltanthura as they share the triangular pleotelson and tapering uropodal exopod, but we refrain from transferring it to Deltanthura as its description lacks the other diagnostic characters of Deltanthura. We provide a revised key to all genera in Paranthuridae Menzies & Glynn, 1968.
  • The protozoan Lagenophrys sp. (Peritrichida: Sessilida: Lagenophryidae) symbiotic on Cryptocandona sp. (Ostracoda: Podocopa: Candonidae) from Hokkaido, Japan.
    Keiichi Kakui; Mizuho Munakata
    Bulletin of the Otaru Museum, 35, 13, 16, 20 Mar. 2022, [Lead author, Corresponding author]
    Japanese, Research institution
  • Predation of tanaidacean crustaceans by Haliclystus tenuis (Cnidaria: Staurozoa) in captivity
    Keiichi Kakui
    Bulletin of the Otaru Museum, 35, 9, 12, 20 Mar. 2022, [Lead author, Corresponding author]
    Japanese, Research institution
  • Three Free-Living Marine Nematodes from Sagami Bay, Japan, with a Description of Wiesoncholaimus jambio sp. nov. (Nematoda: Oncholaimidae)
    Daisuke Shimada; Keiichi Kakui
    Zoological Science, 39, 1, 106, 114, Zoological Society of Japan, 01 Feb. 2022, [Peer-reviewed], [Last author]
    English, Scientific journal
  • Description of a New Hamatipeda Species, with an 18S Molecular Phylogeny (Crustacea: Tanaidacea: Typhlotanaidae)
    Keiichi Kakui; Chizue Hiruta
    Zoological Science, 39, 1, 140, 146, Zoological Society of Japan, 01 Feb. 2022, [Peer-reviewed], [Lead author, Corresponding author], [Domestic magazines]
    English, Scientific journal, We describe a new typhlotanaid species, Hamatipeda kohtsukai sp. nov., collected from between 167 and 488 m depth in the Sagami Sea, Japan. This is the first record of Hamatipeda from the northern hemisphere. Hamatipeda kohtsukai resembles Hamatipeda trapezoida from the Subantarctic region in having pereonites 1-3 widest anteriorly (not rectangular), but differs from it in the length ratio of antennal articles 4/5; the number of setae on the dactyli of pereopods 1-3, ischia of pereopods 4-6, and carpi of pereopods 4-6; the shape of the unguis of pereopods 4-6; and the shape of the uropodal endopod. We determined partial sequences for the cytochrome c oxidase subunit I (COI; cox1) and 18S rRNA (18S) genes in H. kohtsukai. A phylogenetic reconstruction based on the 18S sequences recovered a highly supported Typhlotanaidae clade containing H. kohtsukai and Typhlotanais mixtus, with Paranarthrura sp. (Agathotanaidae) as the sister taxon. A key to species of Hamatipeda is presented., 13264296
  • First record of Scutariella japonica (Platyhelminthes: Rhabdocoela) from Hokkaido, Japan, and notes on its host shrimp Neocaridina sp. aff. davidi (Decapoda: Caridea: Atyidae)
    Keiichi Kakui; Tomoyuki Komai
    Aquatic Animals, 2022, AA2022-1, Jan. 2022, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Comprehensive Transcriptome Sequencing of Tanaidacea with Proteomic Evidences for Their Silk
    Keiichi Kakui; James F Fleming; Masaru Mori; Yoshihiro Fujiwara; Kazuharu Arakawa
    Genome Biology and Evolution, 13, 12, evab281, Oxford University Press (OUP), Dec. 2021, [Peer-reviewed], [Lead author], [International Magazine]
    English, Scientific journal, Abstract
    Tanaidaceans are small benthic crustaceans that mainly inhabit diverse marine environments, and they comprise one of the most diverse and abundant macrofaunal groups in the deep sea. Tanaidacea is one of the most thread-dependent taxa in the Crustacea, constructing tubes, spun with their silk, for shelter. In this work, we sequenced and assembled the comprehensive transcriptome of 23 tanaidaceans encompassing 14 families and 4 superfamilies of Tanaidacea, and performed silk proteomics of Zeuxo ezoensis to search for its silk genes. As a result, we identified two families of silk proteins that are conserved across the four superfamilies. The long and repetitive nature of these silk genes resembles that of other silk-producing organisms, and the two families of proteins are similar in composition to silkworm and caddisworm fibroins, respectively. Moreover, the amino acid composition of the repetitive motifs of tanaidacean silk tends to be more hydrophilic, and therefore could be a useful resource in studying their unique adaptation of silk use in a marine environment. The availability of comprehensive transcriptome data in these taxa, coupled with proteomic evidence of their silk genes, will facilitate evolutionary and ecological studies., 10432430;11341431;13264296;10432431
  • A new species of Expanathura (Crustacea: Isopoda: Anthuroidea) from Iriomote Island, Japan, with a note on male polymorphism
    SHOKI SHIRAKI; MICHITAKA SHIMOMURA; KEIICHI KAKUI
    Zootaxa, 5047, 3, 377, 390, Magnolia Press, 30 Sep. 2021, [Peer-reviewed], [Last author]
    English, Scientific journal, We describe Expanathura monile sp. nov. from Iriomote Island, Okinawa, Japan, northwestern Pacific. This species resembles E. collaris, E. macronesia, and E. haddae in having very broad uropodal rami, a broad telson, and a moderately shortened pleopod 1 endopod, but differs from these three species in the female having (1) the head with a dorsal V-shaped band of brown pigmentation, (2) the antennular flagellum with 3–4 articles, (3) antennal peduncular article 2 with an outer triangular projection, (4) the uropodal endopod longer than wide, (5) the uropodal exopod with an acute triangular projection, (6) an oval telson, and (7) the posterolateral setae on the telson similar in length. We present a revised key to known species in the genus Expanathura and briefly discuss the male polymorphism observed in E. monile. , 11341430
  • A new cave-dwelling genus and species of Nerillidae (Annelida) from the Ryukyu Islands, Japan
    Katrine Worsaae; Malte J. Hansen; Oscar Axelsen; Keiichi Kakui; Peter R. Møller; Karen J. Osborn; Alejandro Martínez; Brett C. Gonzalez; Norio Miyamoto; Yoshihisa Fujita
    Marine Biodiversity, 51, 4, Springer Science and Business Media LLC, 21 Jul. 2021, [Peer-reviewed]
    English, Scientific journal
  • Mismatch between similarity of mitochondrial gene order and phylogenetic distance in Podocopa (Crustacea: Ostracoda)
    Keiichi Kakui; Mizuho Munakata; Hayato Tanaka; Chizue Hiruta
    Zoologischer Anzeiger, 293, 145, 148, Elsevier BV, Jul. 2021, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • First report of marine horsehair worms (Nematomorpha: Nectonema) parasitic in isopod crustaceans
    Keiichi Kakui; Jun Fukuchi; Daisuke Shimada
    Parasitology Research, 120, 7, 2357, 2362, Springer Science and Business Media LLC, Jul. 2021, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Presumptive stridulatory organs in Paranthura cf. japonica Richardson, 1909 (Isopoda: Cymothoida: Paranthuridae)
    Keiichi Kakui; Shoki Shiraki
    Journal of Crustacean Biology, 41, 2, Oxford University Press ({OUP}), 11 Jun. 2021, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 13264296
  • First Complete Mitochondrial Genome of a Tanaidacean Crustacean (Arctotanais alascensis)
    Keiichi Kakui; Yasunori Kano
    Zoological Science, 38, 3, 267, 272, Zoological Society of Japan, 01 Jun. 2021, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 13264296
  • Heterocypris spadix sp. nov. (Crustacea: Ostracoda: Cypridoidea) from Japan, with Information on Its Reproductive Mode
    Mizuho Munakata; Hayato Tanaka; Keiichi Kakui
    Zoological Science, 38, 3, 287, 296, Zoological Society of Japan, 01 Jun. 2021, [Peer-reviewed], [Last author]
    English, Scientific journal, 11341431
  • Two New Species of Free-living Marine Nematodes (Nematoda: Axonolaimidae and Tripyloididae) from the Coast of Antarctica
    Daisuke Shimada; Atsushi C. Suzuki; Megumu Tsujimoto; Satoshi Imura; Keiichi Kakui
    Species Diversity, 26, 1, 49, 63, Japanese Society of Systematic Zoology, 22 Mar. 2021, [Peer-reviewed], [Last author]
    English, Scientific journal
  • Zeuxo ezoensis (Crustacea: Tanaidacea) from an open-air pool on Rishiri Island
    Keiichi Kakui; Mizuho Munakata; Shinri Tomioka
    Rishiri Studies, 40, 21, 24, 01 Mar. 2021, [Lead author, Corresponding author]
    Japanese, Research institution
  • Tube-constructing paratanaoidean tanaidaceans (Crustacea: Peracarida): a brief review
    Keiichi Kakui
    Aquatic Animals, AA2021, AA2021-4, 12 Feb. 2021, [Peer-reviewed], [Domestic magazines]
    English, Scientific journal, 13264296
  • Dirivultidae (Copepoda: Siphonostomatoida) from hydrothermal vent fields in the Okinawa Trough, North Pacific Ocean, with description of one new species
    Daisuke Uyeno; Keiichi Kakui; Hiromi K. Watanabe; Yoshihiro Fujiwara
    Journal of the Marine Biological Association of the United Kingdom, 100, 8, 1283, 1298, Cambridge University Press ({CUP}), 10 Feb. 2021, [Peer-reviewed]
    English, Scientific journal, AbstractTwo species of dirivultid copepod (Siphonostomatoida), Stygiopontius senckenbergi Ivanenko & Ferrari, 2013 and Dirivultus kaiko sp. nov., were discovered from hydrothermal vent fields in the Okinawa Trough, the western North Pacific. Since S. senckenbergi was originally described based on two adult males from the New Ireland Fore-Arc system, Papua New Guinea, the discovery here represents the second record. This species was attached on ventral setae covered with filamentous bacteria of the deep-sea squat lobster Shinkaia crosnieri (Decapoda: Munidopsidae). The female of S. senckenbergi is described for the first time as well as the nauplius I and the copepodid IV. Sexual dimorphism is shown on several appendages (i.e. antennule, leg 2 and leg 5). The nauplius I shows typical lethithotrophic characteristics. Dirivultus kaiko sp. nov., which was found on the tentacular crown of the siboglinid tubeworm Lamellibrachia columna, differs from its two congeners by the antennule, maxilliped and leg 5 of both sexes. These are the second and third dirivultid species to be reported from Japan., 11341431
  • A New Species of Free-living Marine Nematode, Proplatycoma tsukubae sp. nov. (Enoplida: Leptosomatidae), from Japan
    Daisuke Shimada; Keiichi Kakui
    Species Diversity, 25, 2, 205, 211, Japanese Society of Systematic Zoology, 30 Nov. 2020, [Peer-reviewed], [Last author]
    English, Scientific journal, 11341431
  • Unexpected low genetic differentiation between Japan and Bering Sea populations of a deep-sea benthic crustacean lacking a planktonic larval stage (Peracarida: Tanaidacea)
    Keiichi Kakui; Hidetaka Nomaki; Hironori Komatsu; Yoshihiro Fujiwara
    Biological Journal of the Linnean Society, 131, 3, 566, 574, Oxford University Press ({OUP}), 06 Nov. 2020, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, AbstractInformation on the extent, diversity and connectivity of populations is lacking for most deep-sea invertebrates. Species of the order Tanaidacea (Crustacea), one of the most diverse and abundant macrofaunal groups in the deep sea, are benthic, lack a planktonic larval stage, and thus would be expected to have narrow distributional ranges. However, with molecular evidence from the COI gene, we show here that the deep-sea tanaidacean Carpoapseudes spinigena has a distributional range spanning at least 3700 km, from off northern Japan to the south-eastern Bering Sea. Living individuals found in a sediment core indicated that the species is a sedentary burrower. COI analyses revealed a low level of genetic diversity overall, and low differentiation (p-distance, 0.2–0.8%) between the Japan and Bering Sea populations. One hypothesis to explain the low genetic diversity over a broad region is that the Japan population was founded by individuals transported by ocean currents from the Bering Sea. However, due to limited data, other explanations cannot be ruled out. Our results indicate that continued sampling is of fundamental importance to understanding how genetic and taxonomic diversity originate and are maintained in the deep sea., 11341431
  • A new species of Zeuxo (Crustacea: Peracarida: Tanaidacea) from Japan, with remarks on carapace pigmentation as a potentially useful taxonomic character
    Nobuya Okamoto; Yuki Oya; Keiichi Kakui
    Marine Biology Research, 16, 5, 411, 422, Informa UK Limited, 22 Oct. 2020, [Peer-reviewed], [Last author]
    English, Scientific journal, 11341431;13264296
  • A new species of Pantopipetta (Pycnogonida: Austrodecidae) from the North Pacific, with a note on the palp articulation
    Yushi Hosoda; Keiichi Kakui
    Marine Biology Research, 16, 5, 390, 395, Informa UK Limited, 22 Oct. 2020, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal
  • New replacement name for a homonym in Lepeophtheirus (Crustacea: Copepoda: Caligidae)
    Kakui K; Uyeno D
    Zootaxa, 4852, 1, 143, 144, Magnolia Press, 14 Sep. 2020, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, Markevich (1940) established Pseudolepeophtheirus Markevich, 1940 for Pseudolepeophtheirus longicauda Markevich, 1940 based on copepods collected from the pleuronectid fish Platichthys stellatus (Pallas, 1787). Dojiri & Ho (2013) synonymized the genus and the species with Lepeophtheirus Nordmann, 1832 and Lepeophtheirus parvicruris Fraser, 1920, respectively. Later, Homma et al. (2020) resurrected Markevich’s species as a member of Lepeophtheirus, i.e., as L. longicauda (Markevich, 1940). The last component of the names of both genera is ‘phtheirus’ (transliterated from the Greek φθειρ; Nordmann 1832: 30), a masculine noun, and thus under Article 30.1.2 of the International Code of Zoological Nomenclature (hereinafter, Code; International Commission on Zoological Nomenclature 1999), both generic names are also masculine. The species-group name longicauda might be regarded as either a noun in apposition or as an adjective in the feminine gender, and Markevich (1940) did not specify his intention in this regard. Bearing in mind that ‘cauda’, meaning ‘tail’, actually is a feminine Latin noun and that Markevich did not change the final ‘-a’ to ‘-us’ to match the masculine gender of the genus, we deem that longicauda Markevich, 1940 is a noun in apposition, a position supported by Article 31.2.2 of the Code.
  • A new species of Nicippe from the Bering Sea (Crustacea, Amphipoda, Pardaliscidae), with a redescription of N. tumida
    Ko Tomikawa; Keiichi Kakui; Yoshihiro Fujiwara
    ZooKeys, 965, 965, 37, 53, 03 Sep. 2020, [Peer-reviewed]
    English, Scientific journal, 11341431
  • First in situ observations of behavior in deep-sea tanaidacean crustaceans
    Kakui K; Fujiwara Y
    Zoological Science, 37, 4, 303, 306, 03 Aug. 2020, [Peer-reviewed], [Lead author, Corresponding author], [Domestic magazines]
    English, Scientific journal, We report on the behavior of a deep-sea tanaidacean, Gigantapseudes sp. (Apseudomorpha: Gigantapseudidae), recorded at the depths of 6446-6447 m by the manned submersible Shinkai 6500. From recordings of at least three individuals walking on the sea floor, we confirm that Gigantapseudes sp. is epibenthic, as previously inferred from leg shape. One individual was recorded entering a hole. All individuals in the videos kept pereopods 4 raised from the seafloor while walking, implying that those legs have a function other than for walking, such as mechano- or chemoreception, or posture control. Our in situ observations of behavior are the first for any deep-sea tanaidacean and illustrate the importance of recording high-resolution videos in the deep sea and archiving them for future use. Our identification of Gigantapseudes sp. from video footage provides the first record of this genus from Japanese waters and extends the northern limit of the known generic distribution.
  • Integrative taxonomy of Pseudolepeophtheirus longicauda (Crustacea: Copepoda: Caligidae) parasitic on Platichthys stellatus (Actinopterygii: Pleuronectidae)
    Homma R; Uyeno D; Kakui K
    Parasitology International, 78, 102135, 24 Jul. 2020, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal
  • First record of a tanaidacean crustacean fed upon by an arrow worm (Chaetognatha)
    Kakui K
    Bulletin of the Kitakyushu Museum of Natural History and Human History Series A (Natural History), 18, 1, 4, 31 Mar. 2020, [Peer-reviewed]
    English, Scientific journal, 13264296
  • Paradoxapseudes shimojiensis sp. nov. (Crustacea: Tanaidacea: Apseudidae) from a submarine limestone cave in Japan, with notes on its chelipedal morphology and sexual system
    Kakui K; Fujita Y
    Marine Biology Research, 16, 3, 195, 207, Informa {UK} Limited, 15 Mar. 2020, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 11341431;13264296
  • First record of Conchoderma auritum (Linnaeus, 1767) (Crustacea: Cirripedia: Lepadidae) from Rishiri Island
    Tomioka S; Kakui K; Kado R; Sato M
    Rishiri Studies, 39, 7, 10, Mar. 2020
    Japanese, Research institution
  • Shell-exchange behavior in a hermit-crab-like tanaidacean (Crustacea: Malacostraca)
    Kakui K
    Zoological Science, 36, 468, 470, Dec. 2019, [Peer-reviewed]
    English, Scientific journal
  • First report of Teleotanaidae (Crustacea: Tanaidacea) from the Northwestern Pacific, with the description of a new species
    Tanabe Y; Kakui K
    Species Diversity, 24, 181, 187, Nov. 2019, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal, 12050400;11341431
  • Benthic deep-sea fauna in south of the Kii Strait and the Sea of Kumano, Japan. Second report
    Kimura Taeko; Kimura Shoichi; Kakui Keiichi; Hookabe Natsumi; Kuramochi Toshiaki; Fujita Toshihiko; Ogawa Akito; Kobayashi Itaru; Jimi Naoto; Okanishi Masanori; Yamaguchi Haruka; Hirose Masato; Yoshikawa Akihiro; Fukuchi Jun; Shimomura Michitaka; Kashio Sho; Uyeno Daisuke; Fujiwara Kyoji; Naruse Tohru; Kushida Yuka; Kise Hiroki; Maekawa Yoichi; Nakamura Toru; Okumura Junya; Tanaka Kazuki
    Annals of Field Research and Technology Mie University, 17, 17, 1, 29, 三重大学大学院生物資源学研究科, Nov. 2019
    Japanese, Scientific journal, Preliminary results of the deep-sea faunal survey conducted from the TR/V Seisui-maru of Mie University in April 2019 are presented. A total of 20 taxonomists and ecologists working on a wide variety of animal taxa participated in this survey. Surveyed areas included the Kumano Sea(off Mie Prefecture)and south of the Kii Strait(off Tanabe Bay, Wakayama Prefecture), at depths of 112-775 m. Sampling gears employed were beam trawl and biological dredge. The collection is represented by macrobenthos and meiobenthos from nine animal phyla, including echinoderms, arthropods, molluscans and annelids. The number of phyla occurring in each station varied from seven to eight. The station with most diverse fauna at the phylum level was St. 3B(south of the Kii Strait, 775-661 m depth, mud bottom).Meiofauna includes priapulids, nematodes and small arthropods such as copepods, tanaidaceans, amphipods, isopods and cumaceans. In addition to freeliving species, cnidarians symbiotic on a gastropod and an antipatharian, and crustaceans parasitic on a fi sh and sea urchins were also collected. Preliminary identifi cations are given for Asteroidea, Ophiuroidea, Holothuroidea, Crinoidea, Cyclopoida, Siphonostomatoida, Tanaidacea, Isopoda, Decapoda, Mollusca, polychaetes, Bryozoa, Cnidaria and Nemertea., 13264296
  • On the authorship of the genus-group name Curtipleon (Crustacea: Tanaidacea: Metapseudidae)
    Kakui K; Nakano T
    Species Diversity, 24, 179, 180, Nov. 2019, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Sexual system in the tanaidacean Falsapseudes bowmani (Crustacea: Malacostraca: Peracarida)
    Kakui K; Hiruta C; Uyeno D
    Invertebrate Biology, 138, 3, e12257, Wiley, Sep. 2019, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Two Hexapleomera species from Japan, with a new species description and discussion of phylogenetic relationships within Hexapleomera (Crustacea: Tanaidacea)
    Tanabe Y; Kakui K
    Zootaxa, 4648, 318, 336, Jul. 2019, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal, We describe Hexapleomera sasuke sp. nov. and provide a supplemental redescription of Hexapleomera urashima Tanabe et al., both of which are based on specimens collected in Japan. Hexapleomera sasuke closely resembles H. urashima, sharing a uropod with four articles, maxillipedal endites with two tiny dorso-subdistal and two distal spiniform setae, a labium with the palp fused to the outer lobe, and the pereopod-1 propodus with an inner subdistal plumose seta. The new species differs from H. urashima in having the basal article of pleopod 3 with three outer plumose setae, the chelipedal carpus with three or four dorsodistal simple setae, the region between the bases of the chelipedal dactylus and fixed finger with three or four simple setae, and deeply pigmented pleopodal rami. We determined partial sequences of the cytochrome c oxidase subunit I (COI) gene (655 nt) from H. sasuke, which proved to be 15.0–15.3% divergent from H. urashima in Kimura 2-parameter (K2P) distance; also in partial sequences of 18S rRNA gene (1888 nt after alignment), 0.8% divergent (K2P distance) was detected between two species. Based on morphology and molecular data, we discuss phylogenetic relationships within Hexapleomera, and propose two morphologically distinct groups, the robusta group and the wombat group., 11341431
  • Two new and one known species of Phanodermatidae (Nematoda: Enoplida) from Sagami Bay, Japan
    Shimada D; Kakui K
    Zootaxa, 4608, 484, 500, May 2019, [Peer-reviewed], [Last author]
    English, Scientific journal, Two new and one known species of free-living marine nematode belonging to the family Phanodermatidae are described. They were collected during the 12th Japanese Association for Marine Biology (JAMBIO) Coastal Organism Joint Survey off Jogashima, Japan in Sagami Bay, North Pacific Ocean. Phanodermopsis kohtsukai sp. nov. differs from the congeners by the shape of the tail, the longer spicules, and the absence of the gubernaculum. Micoletzkyia nakanoi sp. nov. differs from the congeners by the absence of a circle of cervical setae, the proportion of the tail, the cylindrical gubernaculum without projection, and the position of the vulva. Crenopharynx caudata (Filipjev, 1927) comb. nov. is transferred from the genus Phanodermopsis based on the presence of the long spicules. The morphological characters of the newly collected specimens of C. caudata comb. nov. are in excellent agreement with those shown in the original description of the specimens from Arctic Russia and the previous redescription of the specimens from Skagerrak. The new diagnoses and species lists for these three genera and two dichotomous keys to the males of Micoletzkyia and Crenopharynx are provided. 
  • Topotype-based DNA barcode of the parasitic isopod Pseudione nephropsi (Bopyridae), with a supplementary morphological description
    Kakui K; Shimomura M; Kimura S; Kimura T
    Species Diversity, 24, 103, 108, May 2019, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 11341431
  • First molecularly confirmed conspecific male and female pair in Konariinae (Crustacea: Tanaidacea), with the description of a new species
    Kakui K; Uyeno D; Naruse T
    Species Diversity, 24, 1, 49, 60, Japanese Society of Systematic Zoology, May 2019, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 11341431
  • First record of Hemichela nanhaiensis (Pycnogonida: Ammotheidae) from Japanese waters, with the first description of females
    Hosoda Y; Tomioka S; Kakui K
    Species Diversity, 23, 215, 218, Nov. 2018, [Peer-reviewed], [Last author]
    English, Scientific journal, 10432430
  • Benthic deep-sea fauna in the Sea of Kumano, Mie Prefecture, Japan
    Kimura Taeko; Kimura Shoichi; Jimi Naoto; Kakui Keiichi; Tomioka Shinri; Oya Yuki; Matsumoto Yu; Tanabe Yuki; Hasegawa Naohiro; Hookabe Natsumi; Homma Riko; Hosoda Yushi; Fujimoto Shinta; Kuramochi Toshiaki; Fujita Toshihiko; Ogawa Akito; Kobayashi Itaru; Ishida Yoshiaki; Tanaka Hayate; Onishi Haruka; Shimetsugu Miho; Yoshikawa Akihiro; Tanaka Masaatsu; Kushida Yuka; Maekawa Yoichi; Nakamura Toru; Okumura Junya; Tanaka Kazuki
    Annals of Field Research and Technology Mie University, 16, 16, 1, 32, 三重大学大学院生物資源学研究科, Oct. 2018
    Japanese, Research institution, In this study, we had investigated the benthic deep-sea fauna using the dredge and beam trawl in the region from the continental shelf to the continental slope of the Sea of Kumano, Mie Prefecture, during the No.1722 research voyage of the training ship Seisui-Maru of Mie University. The survey was carried out at 16 stations covering a depth range of 113-1059 m. The results of the survey, 14 phyla had been confirmed. Arthropod, echinoderm, annelid and molluscan macrobenthos were collected from all of the stations. The phylum number of each station was in the range from 4 to 11. The largest number of phyla had been confi rmed at St.10D of boulders bottom(768-800 m depth). Meiobenthos confi rmed in our sample were kinorhynchs, nematodes, tardigrades, loriciferans and small arthropods such as tanaidaceans, copepods and cumaceans. In addition to free-living species, parasitic copepods, isopods, platyhelminthes, acanthocephalans and nematodes had been found in fish, crustaceans and polychaetes., 11341431
  • Molecular Phylogeny of the Family Capitellidae (Annelida)
    Shinri Tomioka; Keiichi Kakui; Hiroshi Kajihara
    ZOOLOGICAL SCIENCE, 35, 5, 436, 445, Oct. 2018, [Peer-reviewed]
    English, Scientific journal
  • First report of the chelonophilic tanaidid Hexapleomera urashima from an environment other than the surface of sea turtles
    Keiichi Kakui
    Fauna Ryukyuana, 45, 5, 8, Sep. 2018, [Peer-reviewed], [Lead author, Corresponding author], [Domestic magazines]
    English, Scientific journal, 10905851;11341431
  • Targeted gene disruption by use of CRISPR/Cas9 ribonucleoprotein complexes in the water flea Daphnia pulex
    Chizue Hiruta; Keiichi Kakui; Knut E. Tollefsen; Taisen Iguchi
    Genes to Cells, 23, 6, 494, 502, Blackwell Publishing Ltd, 01 Jun. 2018, [Peer-reviewed]
    English, Scientific journal
  • Tsuranarthrura shinsei, a new genus and species in Anarthruridae (Crustacea: Tanaidacea) from the Northwestern Pacific
    Keiichi Kakui; Shinri Tomioka
    Species Diversity, 23, 1, 61, 68, Japanese Society of Systematic Zoology, 2018, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 11341431
  • Haimormus shimojiensis, a new genus and species of Pseudozeuxidae (Crustacea: Tanaidacea) from a submarine limestone cave in Northwestern Pacific
    Keiichi Kakui; Yoshihisa Fujita
    PeerJ, 2018, 4, e4720, PeerJ Inc., 2018, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 11341431
  • A new species of Xenoturbella from the western Pacific Ocean and the evolution of Xenoturbella
    Hiroaki Nakano; Hideyuki Miyazawa; Akiteru Maeno; Toshihiko Shiroishi; Keiichi Kakui; Ryo Koyanagi; Miyuki Kanda; Noriyuki Satoh; Akihito Omori; Hisanori Kohtsuka
    BMC Evolutionary Biology, 18, 1, BioMed Central Ltd., 18 Dec. 2017, [Peer-reviewed]
    English, Scientific journal
  • A new species of Xenoturbella from the western Pacific Ocean and the evolution of Xenoturbella
    Hiroaki Nakano; Hideyuki Miyazawa; Akiteru Maeno; Toshihiko Shiroishi; Keiichi Kakui; Ryo Koyanagi; Miyuki Kanda; Noriyuki Satoh; Akihito Omori; Hisanori Kohtsuka
    BMC EVOLUTIONARY BIOLOGY, 17, 1, 245, Dec. 2017, [Peer-reviewed]
    English, Scientific journal
  • Oncholaimus langhovdensis sp. nov. (Nematoda: Enoplea: Oncholaimida), a new species of free-living marine nematode from Langhovde, Dronning Maud Land, East Antarctica
    Daisuke Shimada; Atsushi C. Suzuki; Megumu Tsujimoto; Satoshi Imura; Keiichi Kakui
    Species Diversity, 22, 2, 151, 159, Japanese Society of Systematic Zoology, 01 Dec. 2017, [Peer-reviewed], [Last author]
    English, Scientific journal
  • Hexapleomera urashima sp nov (Crustacea: Tanaidacea), a tanaidid epibiotic on loggerhead sea turtles at Yakushima Island, Japan
    Yuki Tanabe; Ryota Hayashi; Shinri Tomioka; Keiichi Kakui
    ZOOTAXA, 4353, 1, 146, 160, Nov. 2017, [Peer-reviewed], [Last author]
    English, Scientific journal, 11341431
  • Tube construction by a tanaidacean crustacean using a novel mucus secretion system involving the anal opening
    Keiichi Kakui; Chizue Hiruta
    ZOOLOGICAL LETTERS, 3, Nov. 2017, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 11341431
  • Morphology of a newly recorded species, Eremaeozetes octomaculatus Hammer, 1973 from Indonesia (Acari: Oribatida: Eremaeozetidae), with its genetic information
    Badamdorj Bayartogtokh; Satoshi Shimano; Wataru Hagino; Keiichi Kakui; Shimpei F. Hiruta; Sri Hartini
    International Journal of Acarology, 43, 6, 444, 449, Taylor and Francis Inc., 18 Aug. 2017, [Peer-reviewed]
    English, Scientific journal
  • A new species of Tanaopsis (Crustacea: Tanaidacea) from Japan, with remarks on the functions of serial ridges and grooves on the appendages
    Keiichi Kakui; Daisuke Shimada
    ZOOTAXA, 4282, 2, 324, 336, Jun. 2017, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, 11341431
  • Difference in Size at Maturity in Annual and Overwintering Generations in the Tanaidacean Zeuxo sp in Oshoro Bay, Hokkaido, Japan
    Keiichi Kakui; Yoshinobu Hayakawa; Haruo Katakura
    ZOOLOGICAL SCIENCE, 34, 2, 129, 136, Apr. 2017, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Habitat of a tanaidacean Apseudes nipponicus Shiino, 1937
    Kakui K; Suzuki A; Nakano H; Kohtsuka H
    Bulletin of the Kitakyushu Museum of Natural History and Human History Series A (Natural History), 15, 1, 3, Mar. 2017, [Peer-reviewed], [Lead author, Corresponding author]
    English, Research institution
  • Rediscovery and redescription of the abyssal squat lobster Munidopsis petalorhyncha Baba, 2005 (Crustacea: Decapoda: Munidopsidae) from the Northwest Pacific
    Tomoyuki Komai; Ivan Marin; Keiichi Kakui
    ZOOTAXA, 4226, 1, 93, 102, Jan. 2017, [Peer-reviewed], [Last author]
    English, Scientific journal
  • A new species and the shallowest record of Flabegraviera Salazar-Vallejo, 2012 (Annelida: Flabelligeridae) from Antarctica
    Naoto Jimi; Megumu Tsujimoto; Kentaro Watanabe; Keiichi Kakui; Hiroshi Kajihara
    ZOOTAXA, 4221, 4, 477, 485, Jan. 2017, [Peer-reviewed]
    English, Scientific journal
  • Review of the taxonomy, diversity, ecology, and other biological aspects of Order Tanaidacea from Japan and surrounding waters
    Kakui K
    In: Motokawa M, Kajihara H (Eds) Species Diversity of Animals in Japan, Springer, 603, 627, Dec. 2016, [Peer-reviewed], [Invited]
    English, Scientific journal
  • An overview of recent marine-biodiversity research in Japan
    Kajihara H; Kakui K
    In: Motokawa M, Kajihara H (Eds) Species Diversity of Animals in Japan, Springer, 25, 45, Springer Japan, Dec. 2016, [Peer-reviewed], [Invited], [Last author]
    English, Scientific journal
  • Opisa takafuminakanoi, a new species of Opisidae from Hokkaido, Japan (Crustacea: Amphipoda)
    Yukiko Narahara-Nakano; Keiichi Kakui; Ko Tomikawa
    ZOOTAXA, 4200, 2, 335, 339, Nov. 2016, [Peer-reviewed]
    English, Scientific journal
  • Cosmopolitan or Cryptic Species? A Case Study of Capitella teleta (Annelida: Capitellidae)
    Shinri Tomioka; Tomohiko Kondoh; Waka Sato-Okoshi; Katsutoshi Ito; Keiichi Kakui; Hiroshi Kajihara
    ZOOLOGICAL SCIENCE, 33, 5, 545, 554, Oct. 2016, [Peer-reviewed]
    English, Scientific journal
  • Three new brackish-water thalassocypridine species (Crustacea: Ostracoda: Paracyprididae) from the Ryukyu Islands, southwestern Japan
    Shimpei F. Hiruta; Keiichi Kakui
    ZOOTAXA, 4169, 3, 515, 539, Sep. 2016, [Peer-reviewed], [Last author]
    English, Scientific journal
  • Diversity of sexual and reproductive systems in Tanaidacea
    Kakui K
    Cancer, 25, 25, 131, 136, 日本甲殻類学会, Aug. 2016, [Invited]
    Japanese, Scientific journal
  • Multiple processes generate productivity-diversity relationships in experimental wood-fall communities
    Craig R. McClain; James P. Barry; Douglas Eernisse; Tammy Horton; Jenna Judge; Keiichi Kakui; Chris Mah; Anders Waren
    ECOLOGY, 97, 4, 885, 898, Apr. 2016, [Peer-reviewed]
    English, Scientific journal
  • Mesoscale morphology at nanoscale resolution: serial block-face scanning electron microscopy reveals fine 3D detail of a novel silk spinneret system in a tube-building tanaid crustacean
    Tomonari Kaji; Keiichi Kakui; Naoyuki Miyazaki; Kazuyoshi Murata; A. Richard Palmer
    FRONTIERS IN ZOOLOGY, 13, Mar. 2016, [Peer-reviewed]
    English, Scientific journal
  • Tubulanus tamias sp nov (Nemertea: Palaeonemertea) with Two Different Types of Epidermal Eyes
    Hiroshi Kajihara; Keiichi Kakui; Hiroshi Yamasaki; Shimpei F. Hiruta
    ZOOLOGICAL SCIENCE, 32, 6, 596, 604, Dec. 2015, [Peer-reviewed]
    English, Scientific journal
  • The first record of a species of Clausiidae (Copepoda: Cyclopoida) from Japanese waters, with the proposal of a new genus.
    Uyeno D; Kakui K
    Species Diversity, 20, 2, 153, 158, Japanese Society of Systematic Zoology, Nov. 2015, [Peer-reviewed], [Last author]
    English, Scientific journal
  • JAMBIO Coastal Organism Joint Surveys reveals undiscovered biodiversity around Sagami Bay
    Hiroaki Nakano; Keiichi Kakui; Hiroshi Kajihara; Michitaka Shimomura; Naoto Jimi; Shinri Tomioka; Hayate Tanaka; Hiroshi Yamasaki; Masaatsu Tanaka; Takato Izumi; Masanori Okanishi; Yutaro Yamada; Hideo Shinagawa; Toshihiko Sato; Yasutaka Tsuchiya; Akihito Omori; Mamoru Sekifuji; Hisanori Kohtsuka
    REGIONAL STUDIES IN MARINE SCIENCE, 2, 77, 81, Nov. 2015, [Peer-reviewed]
    English, Scientific journal
  • Indoapseudes bamberi sp nov (Crustacea: Tanaidacea: Pagurapseudidae: Hodometricinae) from Iriomote Island, Ryukyu Islands, southwestern Japan
    Keiichi Kakui; Tohru Naruse
    ZOOTAXA, 3995, 1, 138, 148, Aug. 2015, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • First record of the free-living marine nematode Deontostoma magnificum (Timm, 1951) Platonova, 1962 (Nematoda: Leptosomatidae) from Japan.
    Shimada D; Kakui K
    Nematological Research, 45, 1, 13, 18, Jul. 2015, [Peer-reviewed], [Last author]
    English, Scientific journal
  • Phylogenetic position of the queer, backward-bent entoproct Loxosoma axisadversum (Entoprocta: Solitaria: Loxosomatidae).
    Kajihara H; Tomioka S; Kakui K; Iseto T
    Species Diversity, 20, 1, 83, 88, Japanese Society of Systematic Zoology, May 2015, [Peer-reviewed]
    English, The solitary entoproct Loxosoma axisadversum Konno, 1972 was found on the body surface of the maldanid polychaete Nicomache personata Johnson, 1901 obtained among roots of the seagrass Phyllospadix iwatensis Makino collected at a depth of about 1m in Oshoro Bay, Hokkaido, Japan. The species had previously been known from elsewhere in northern Japan, attached to another maldanid, Nicomache minor Arwidsson, 1907. The morphology of the new material is briefly described with photographs taken in life. A preliminary phylogenetic analysis using concatenated partial sequences of the mitochondrial cytochrome c oxidase subunit I (COI) gene, as well as the nuclear 18S and 28S rRNA genes, placed L. axisadversum as the sister taxon to all the rest of Solitaria included in the analysis, indicating non-monophyly of the genus Loxosoma Keferstein, 1862.
  • Two new shallow-water species of Agathotanais (Crustacea: Tanaidacea) from Japan.
    Kakui K; Kohtsuka H
    Species Diversity, 20, 1, 45, 58, Japanese Society of Systematic Zoology, May 2015, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, We describe two shallow-water species of Agathotanais, A. misakiensis sp. nov. and A. toyoshioae sp. nov., from Japan. Agathotanais misakiensis was collected from 211-493m depth in the Sagami Sea, North Pacific Ocean. It closely resembles A. ghilarovi Kudinova-Pasternak, 1989, but differs in having shorter pereonites 3 and 4, all pleonites narrower than pereonite 6, and a shorter article 3 in the antennule. Agathotanais toyoshioae, collected from 95m depth in the Genkai Sea, Sea of Japan, is very similar to A. spinipoda Larsen, 1999. The former can be distinguished from the latter by its narrower pleonites, the presence of a ventral keel on pleonites 1 and 2, and the presence on the basis and absence on the carpus and propodus of small spines on pereopods 4-6.
  • First report of Zeuxo sp. (Crustacea: Tanaidacea) as prey for the forktongue goby, Chaenogobius annularis Gill, 1859. [in Japanese with English abstract]
    Kakui K
    Rishiri Studies, 34, 1, 6, Mar. 2015
    Japanese, Research institution
  • A novel transmission pathway: first report of a larval trematode in a tanaidacean crustacean.
    Kakui K
    Fauna Ryukyuana, 17, 13, 22, Dec. 2014, [Peer-reviewed]
    English, Scientific journal
  • Northernmost records of Antedon serrata A. H. Clark from Hokkaido. [in Japanese with English abstract]
    Kohtsuka H; Kakui K; Kajihara H
    Bulletin of the Biogeographical Society of Japan, 69, 185, 188, Dec. 2014, [Peer-reviewed]
    Japanese, Scientific journal
  • タナイスの糸.
    角井敬知
    うみうし通信, 84, 2, 3, Sep. 2014, [Invited]
    Japanese, Research institution
  • Diverse Pereopodal Secretory Systems Implicated in Thread Production in an Apseudomorph Tanaidacean Crustacean
    Keiichi Kakui; Chizue Hiruta
    JOURNAL OF MORPHOLOGY, 275, 9, 1041, 1052, Sep. 2014, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Tanaidaceans from Rishiri Island [in Japanese with English Abstract].
    Kakui K; Tomioka S; Yamasaki H
    Rishiri Studies, 33, 7, 12, Mar. 2014
    Japanese, Research institution
  • Tanaidaceans from Rishiri Island (Crustacea: Peracarida)
    Kakui, K.; Tomioka, S.; Yamasaki, H.
    Rishiri Studies, 33, 2014
    Scientific journal
  • Nototanaids (Crustacea: Tanaidacea) from Japan, with the description of a new species of Nototanoides.
    Kakui K; Yamasaki H
    Species Diversity, 2013, 18, 2, 245-254, Dec. 2013, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Description of three species of Halmyrapseudes (Crustacea: Tanaidacea: Parapseudidae), with a discussion of biogeography
    Keiichi Kakui; Saowapa Angsupanich
    ZOOTAXA, 3736, 4, 345, 367, Nov. 2013, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Selfing in a malacostracan crustacean: why a tanaidacean but not decapods
    Keiichi Kakui; Chizue Hiruta
    NATURWISSENSCHAFTEN, 100, 9, 891, 894, Sep. 2013, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • 第166回農林交流センターワークショップ「分子系統学の理論と実習」体験記.
    角井敬知
    日本進化学会ニュース, 14, 1, 23, 25, Mar. 2013, [Invited]
    Japanese, Research society
  • A new species of deep-sea nematode, Micoletzkyia mawatarii sp. nov. (Nematoda: Enoplida: Phanodermatidae) from Northern Japan.
    Shimada D; Kakui K; Kajihara H
    Species Diversity, 17, 2, 221, 226, Dec. 2012, [Peer-reviewed]
    English, Scientific journal
  • A new species of Stegidotea Poore, 1985 (Isopoda: Chaetillidae) from Japan.
    Shimomura M; Kakui K
    Crustaceana Monographs, 17, 303, 314, Sep. 2012, [Peer-reviewed], [Invited], [Last author]
    English, Scientific journal
  • BIRDOTANAIS SONGKHLAENSIS, A NEW GENUS AND SPECIES OF NOTOTANAIDAE (CRUSTACEA: TANAIDACEA) FROM THAILAND
    Keiichi Kakui; Saowapa Angsupanich
    RAFFLES BULLETIN OF ZOOLOGY, 60, 2, 421, 432, Aug. 2012, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • PHYLOGENETIC POSITION OF ARCTOTANAIS IN TRE SUBORDER TANAIDOMORPHA (PERACARIDA: TANAIDACEA)
    Keiichi Kakui; Norio Kobayashi; Hiroshi Kajihara
    JOURNAL OF CRUSTACEAN BIOLOGY, 32, 1, 127, 139, Feb. 2012, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Molecular Systematics of Tanaidacea (Crustacea: Peracarida) Based on 18S Sequence Data, with an Amendment of Suborder/Superfamily-level Classification
    Keiichi Kakui; Toru Katoh; Shimpei F. Hiruta; Norio Kobayashi; Hiroshi Kajihara
    ZOOLOGICAL SCIENCE, 28, 10, 749, 757, Oct. 2011, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • A NEW SPECIES OF THE GENUS INGOLFIELLA HANSEN, 1903 (PERACARIDA, AMPHIPODA) FROM THE OGASAWARA ISLANDS, SOUTHERN JAPAN
    Michitaka Shimomura; Keiichi Kakui
    CRUSTACEANA, 84, 5-6, 543, 551, May 2011, [Peer-reviewed], [Last author]
    English, Scientific journal
  • Pseudosphyrapus cuspidiger sp. nov. from the Nansei Islands, Japan, with a supplementary description of Kudinopasternakia balanorostrata Kakui, Kajihara and Mawatari, 2007 (Crustacea: Tanaidacea: Apseudomorpha: Sphyrapodidae).
    Kakui K; Kajihara H
    Bulletin of the National Museum of Nature and Science, Suppl 5, 53, 70, National Museum of Nature and Science, Feb. 2011, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Digital illustration techniques using graphics software. [in Japanese with English abstract]
    Hiruta SF; Kakui K
    TAXA, Proceedings of the Japanese Society of Systematic Zoology, 29, 29, 19, 30, 日本動物分類学会, Aug. 2010, [Peer-reviewed], [Last author]
    Japanese, Scientific journal, Personal computers (PCs) are now indispensable to any field of study, and it is important to master efficient ways to compose documents, analyze data, and manage literature. This is especially true of illustrations, which take much time and repeated editing prior to use in publications, posters, presentations, and so on. Although some reports exist on techniques for making digital illustrations there has been rapid progress in the development of new PC hardware and software, so that some types of previously intractable digital graphical processing methods are now feasible and even routine. ...
  • A New Species of Psammogammarus (Amphipoda: Melitidae) from Kuchinoerabu Island, Japan, with a Noteon its Feeding Habits
    Ko Tomikawa; Keiichi Kakui; Hiroshi Yamasaki
    ZOOLOGICAL SCIENCE, 27, 7, 615, 626, Jul. 2010, [Peer-reviewed]
    English, Scientific journal
  • A new species of Nesotanais Shiino, 1968 (Crustacea, Tanaidacea) from Japan, with a key to species and a note on male chelipeds
    Keiichi Kakui; Hiroshi Kajihara; Shunsuke F. Mawatari
    ZOOKEYS, 33, 33, 1, 17, 2010, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Two new sphyrapodid species (Crustacea : Tanaidacea : Apseudomorpha) from southwestern Japan
    Keiichi Kakui; Hiroshi Kajihara; Shunsuke F. Mawatari
    ZOOTAXA, 1563, 1563, 37, 54, Aug. 2007, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
■ Other Activities and Achievements
■ Books and other publications
  • 水の中の小さな美しい生き物たち : 小型ベントス・プランクトン百科 : カラー図解
    仲村康秀; 山崎博史; 田中隼人, 節足動物 タナイス類
    朝倉書店, Aug. 2025, 9784254171952, viii, 374p, Japanese, [Contributor]
  • 理科年表 2025
    生物のかたちと系統: 動物の基本型, 菌類の基本型, 生活環, 植物の基本型, 生物の系統(2024年以降監修継続中)
    丸善出版, 28 Nov. 2024, 4621310305, 1190, [Supervisor]
  • プランクトン : クラゲ・ミジンコ・小さな水の生物
    山崎博史; 仲村康秀; 田中隼人, タナイスのなかま
    小学館, Jun. 2024, 9784092172975, 175p, Japanese, [Contributor]
  • 海産無脊椎動物多様性学 100年の歴史とフロンティア
    京都大学フィールド科学教育研究センター瀬戸臨海実験所創立100周年記念出版編集委員会, タナイスの多様性研究
    京都大学学術出版会, Nov. 2022, 9784814004492, [Contributor]
  • 動物学の百科事典
    日本動物学会, 動物界の分類群・系統
    丸善出版, Sep. 2018, 9784621303092, xxv, 770p, 図版 [8] p, Japanese, [Contributor]
  • 平成28年 理科年表 第89冊
    東京天文台; 国立天文台, 動物分類表 甲殻亜門(2015年以降監修継続中)
    東京天文臺, Nov. 2015, 9784621089651, 冊, Japanese, [Supervisor]
■ Syllabus
  • 多様性生物学入門, 2024年, 修士課程, 理学院
  • 多様性生物学研究法, 2024年, 修士課程, 理学院
  • 科学・技術の世界(1単位), 2024年, 学士課程, 全学教育
  • 生物学Ⅱ, 2024年, 学士課程, 全学教育
  • 系統分類学a, 2024年, 学士課程, 理学部
  • 系統分類学Ⅰ, 2024年, 学士課程, 理学部
  • 臨海実習Ⅰ, 2024年, 学士課程, 理学部
  • 臨海実習Ⅰ, 2024年, 学士課程, 理学部
  • ISP生物科学実習Ⅰ・a, 2024年, 学士課程, 理学部
  • ISP生物科学実習Ⅰ・b, 2024年, 学士課程, 理学部
  • 動物系統分類学実習, 2024年, 学士課程, 理学部
  • 動物系統分類学実習, 2024年, 学士課程, 理学部
■ Affiliated academic society
  • May 2025 - Present
    Nippon Cnidaria Ctenophora Biologists
  • Mar. 2025 - Present
    The Systematics Association
  • 2011 - Present
    THE CRUSTACEAN SOCIETY
  • 2008 - Present
    THE CARCINOLOGICAL SOCIETY OF JAPAN
  • 2007 - Present
    THE JAPANESE SOCIETY OF SYSTEMATIC ZOOLOGY
  • 2006 - Present
    THE ZOOLOGICAL SOCIETY OF JAPAN
■ Research Themes
  • 甲殻類に寄生する動物の多様性研究
    2025年度 学術研究助成 自然科学・技術(2)水域生態系保全、循環に関する研究
    Oct. 2025 - Sep. 2026
    公益財団法人クリタ水・環境科学振興財団, Principal investigator, 25B002
  • 網羅的サンプリングに基づく深海底生動物の多様化機構解明
    科学研究費助成事業 基盤研究(B)
    01 Apr. 2022 - 31 Mar. 2026
    狩野 泰則; 遊佐 陽一; 岡西 政典; 角井 敬知
    日本学術振興会, 基盤研究(B), 東京大学, 22H02681
  • Marine and anchialine cave biodiversity in the Ryukyu Islands, Japan
    Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research (B)
    01 Apr. 2020 - 31 Mar. 2025
    藤田 喜久; 下村 通誉; 藤井 琢磨; 井口 亮; 岡西 政典; 角井 敬知; 斉藤 知己
    本研究では、1)洞窟水圏性動物の種多様性、2)洞窟水圏性動物の分子系統・適応進化、3)洞窟水圏における動物群集の構造と個体群維持機構の3つの主要研究テーマに注力した調査研究を実施し、当該地域の洞窟水圏における生物多様性の全容を解明することを目的としている。
    2021年度は、沖縄島と伊江島の海底洞窟における合同野外調査を実施し、さらに徳之島・与論島・与那国島における陸域の洞窟での予備的な野外調査を実施し、2022年度以降の合同調査に備える予定であったが、新型コロナウイルス感染症の影響で調査対象の島々への渡航が厳しく制限される状況に陥り、予定を変更することになった。
    実際、海底洞窟の合同調査は規模を縮小して沖縄島のみで実施し、陸域の洞窟調査は沖永良部島、与論島、多良間島の3カ所に変更した。しかし、初年度(2020年度)及び2021年度にて得られたデータをもとに、以下のような実績を挙げることができた。
    「1)洞窟水圏性動物の種多様性」と「2)洞窟水圏性動物の分子系統・適応進化」については、対象洞窟にて採集された生物の種同定を進め、DNAバーコーディング作業を予定通り進めることができた。
    一方、「3)洞窟水圏における動物群集の構造と個体群維持機構」については、共同研究者が考案した海底洞窟の3D測量手法を沖縄島の海底洞窟にて試験実施した。3D測量図は洞窟形態の特性を可視化するものであり、生物研究を劇的に進展させることが予想されることから、次年度(2022年度)に早急な完成を目指す予定である。また、多良間島において、同島固有の地下水性甲殻類(タラマメアミ)を保護・保全するための生態研究を開始した。
    これらの研究成果(の一部)として、3編の研究論文の出版と、1題の学会発表を実施することができた。また、さらに複数(3編)の論文投稿も行い、これらは2022年度に出版を予定している。
    Japan Society for the Promotion of Science, Grant-in-Aid for Scientific Research (B), Okinawa Prefectural University of Arts, 20H03313
  • 海の底から泳ぎ出す:小型底生甲殻類による泳ぐ矮雄の獲得と進化史
    平成31年度科研費基盤研究(C)
    Apr. 2019 - Mar. 2024
    角井敬知
    日本学術振興会, Principal investigator, Competitive research funding
  • 南西諸島における汽水性タナイス目甲殻類の多様性と進化史
    平成30年度研究助成
    Sep. 2018 - Aug. 2020
    角井敬知
    公益信託ミキモト海洋生態研究助成基金, Principal investigator, Competitive research funding
  • 微小水生甲殻類タナイス目の性表現と生殖様式に関する研究
    2018年度研究助成<奨励>
    Jul. 2018 - Mar. 2019
    角井敬知
    公益財団法人秋山記念生命科学振興財団, Principal investigator, Competitive research funding
  • 深海性タナイス目甲殻類の分類学的研究
    平成30年度共同研究(柏地区外来研究員)
    Apr. 2018 - Mar. 2019
    角井敬知
    東京大学大気海洋研究所, Principal investigator, Competitive research funding
  • 底生動物による海藻葉上固着生活獲得までの進化史の解明
    平成28年度科研費若手研究(B)
    Apr. 2016 - Mar. 2019
    角井敬知
    日本学術振興会, Principal investigator, Competitive research funding
  • 八重山周辺海域に分布する小型甲殻類相のデータベース構築:1.内湾環境
    平成28年度琉球大学共同利用研究
    Apr. 2016 - Mar. 2017
    上野大輔; 下村通誉; 角井敬知
    琉球大学, Principal investigator, Competitive research funding
  • 汽水生小型甲殻類ネソタナイスの分類学的・生殖生物学的研究
    平成26年度(第23回)学術研究助成
    Apr. 2015 - Mar. 2016
    角井敬知
    公益財団法人藤原ナチュラルヒストリー振興財団, Principal investigator, Competitive research funding
  • タナイス目で見つかった,軟甲綱で初となる自家受精種の総合的研究
    平成26年度笹川科学研究助成 学術研究部門
    Apr. 2014 - Mar. 2015
    角井敬知
    公益財団法人日本科学協会, Principal investigator, Competitive research funding
  • 死サンゴ片堆積場(ガレ場)における小型無脊椎動物の多様性調査
    平成26年度琉球大学共同利用研究
    Apr. 2014 - Mar. 2015
    角井敬知
    琉球大学, Principal investigator, Competitive research funding
  • うみさわ会(若手海洋生物研究者フィールドワークショップ)
    2014年度マリンバイオ共同推進機構(JAMBIO)公募型「共同利用・共同研究」
    Apr. 2014 - Mar. 2015
    山崎博史; 角井敬知
    マリンバイオ共同推進機構(JAMBIO), Principal investigator, Competitive research funding
  • タナイス目甲殻類の系統進化学的研究-出糸能力獲得の起源について-
    育成研究助成
    Apr. 2012 - Mar. 2014
    角井敬知
    財団法人水産無脊椎動物研究所, Principal investigator, Competitive research funding
  • タナイス目甲殻類の多様性を生み出した“脚型”の進化史の解明
    特別研究員奨励費
    Apr. 2010 - Mar. 2012
    角井敬知
    本研究はタナイス目甲殻類の多様化に関係する第1歩脚の機能・形態("脚型")の進化史解明を目指すもので,本年度は以下の成果を挙げた.
    タナイス目の分子系統解析の論文を公表した.亜目・上科間の系統関係を初めて明らかにし,得られた系統関係と形態的特徴に基づき分類体系を現行の3亜目3上科から2亜目4上科に変更した.
    所属上科に議論のあったArctotanais alascensisの系統位置を決定,論文として公表した.詳細な形態観察および分子系統解析により,本種がタナイス上科に属することを明らかにした.また,本種に鋏脚座節という,同上科のTanais属と近縁とされるネオタナイス上科のみで報告されていた構造を確認し,2上科の近縁性についてより強い支持を得た.
    アプセウデス亜目の側系統性を示唆する結果を得た.Longiflagrum属とカリアプセウデス科は現在アプセウデス亜目に属しているが,進行中の分子系統解析の結果には,2分類群がタナイス亜目と姉妹群を形成するという結果も含まれた.なお,カリアプセウデス科はタナイス亜目のみで知られる「出糸型」生活も行う可能性が示唆されている科であるため,「出糸型」がアプセウデス亜目に起源するという仮説を立て,組織学的手法も導入した研究を計画中である.
    アプセウデス亜目内で祖先的な脚型と考えられている「俳徊型」を呈する2科のDNAサンプルを作成した.パグラプセウドプシス科は,タイでDr.Angsupanichと共同調査を行い採集した.ギガンタプセウデス科は,琉球海溝から得られた1個体を貸借できた.現在解析を進めているところであるが,前者は貝棲型のグループと姉妹群を形成する派生的なグループであるという結果が得られている.
    なお本年度は上記研究以外に,採集調査中に得た標本に基づき2報の記載論文を共同で執筆し,成果を学会大会で発表した.
    日本学術振興会, 特別研究員奨励費, 北海道大学, Principal investigator, Competitive research funding, 10J01119