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Toshio Kawai
Faculty of Fisheries Sciences　Marine Bioresource and Environmental Science　Marine Biology and Biodiversity
Associate Professor
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Last Updated :2024/05/09

Researcher Profile and Settings

Affiliation

  • Faculty of Fisheries Sciences　Marine Bioresource and Environmental Science　Marine Biology and Biodiversity

Job Title

  • Associate Professor

Research funding number

  • 20462806

J-Global ID

Research Interests

  • 魚類系統分類学、比較解剖学   

Research Areas

  • Life sciences / Biodiversity and systematics
  • Life sciences / Morphology, anatomy

Educational Organization

Academic & Professional Experience

  • 2018/08 - Today Hokkaido University The Hokkaido University Museum
  • 2018/04 - Today Hokkaido University Faculty of Fisheries Sciences
  • 2009/04 - 2018/03 Hokkaido University The Hokkaido University Museum
  • 2007/04 - 2009/03 National Museum of Nature and Science, Tokyo
  • 2006/07 - 2007/03 共和コンクリート工業株式会社 主任研究員

Education

  • 2002/04 - 2006/06  Hokkaido University
  • 2000/04 - 2002/03  Hokkaido University
  • 1996/04 - 2000/03  Hokkaido University  School of Fisheries Sciences

Association Memberships

  • THE JAPANESE SOCIETY OF SYSTEMATIC ZOOLOGY   THE ICHTHYOLOGICAL SOCIETY OF JAPAN   

Research Activities

Published Papers

  • Checklist of fishes collected by bottom otter-trawling from upper Gulf of Thailand by MV-SEAFDEC 2 cruise in December 2019
    V. Vilasri, S. Pattarapongpan, B. Russell, K. Obata, T. Kawai
    The Thailand Natural History Museum Journal 17 (2) 67 - 104 2023/12 [Refereed][Not invited]
  • 北海道函館市臼尻から採集されたモヨウフグArothron stellatus
    小幡光汰, 河合俊郎
    北海道大学水産科学研究彙報 73 (2) 41 - 46 2023/12 [Not refereed][Not invited]
  • Discovery of a second juvenile of the rare flatfish Plagiopsetta glossa (Pleuronectiformes: Samaridae) from the East China Sea
    K. Obata, T. Kawai, G. Shinohara
    Platax 20 67 - 77 2023/12 [Refereed][Not invited]
  • A Rare Flatfish, Samaris spinea (Teleostei: Pleuronectiformes: Samaridae) from the Ogasawara Islands, Japan, with Notes on Its Distribution, Taxonomy and Sexual Dimorphism
    K. Obata, T. Kawai, G. Shinohara
    Bulletin of National Museum of Nature and Science, Ser. A. 49 (4) 179 - 189 2023/11 [Refereed][Not invited]
  • First Thai record of Dasyscopelus brachygnathos (Bleeker, 1856) (Myctophidae) from off Phuket, Andaman Sea
    S. Bangchongmanee, C. Aungtonya, A. Sugiyama, K. Obata, S. Yamashita, T. Kawai
    Phuket Marine Biological Center Research Bulletin 80 101 - 106 2023/10 [Refereed][Not invited]
  • Deep-sea fishes from the Andaman Sea by R/V Chakratong Tongyai during 1996–2000. Part 7: order Stomiiformes
    K. Obata, K. Mita, T. Senda, T. Kawai, F. Tashiro, H. Imamura, C. Aungtonya, S. Banchongmanee
    Phuket Marine Biological Center Research Bulletin 80 21 - 51 2023/10 [Refereed][Not invited]
  • Higuchi Junya, Kim Seong-yong, Kawai Toshio
    Japanese Journal of Ichthyology 一般社団法人 日本魚類学会 advpub 0021-5090 2023 [Refereed][Not invited]
     
    A single specimen of Krusensterniella (Perciformes: Zoarcidae), collected off Tsugaru Peninsula, Aomori, Japan, was identified as Krusensterniella squamosa Chernova, 2022, characterized by three pungent spines in the dorsal fin, 47 dorsal-fin spines, at least 101 dorsal-fin rays and 103 vertebrae, and previously known only from the holotype, collected off Iwate, Japan. The new standard Japanese name “Uroko-haregaji”, referring to the scaled body, is proposed for the species. Records of Krusensterniella notabilis from Japan are unlikely to be that species, since many identification sources for Japanese fishes in correctly described K. notabilis as having a broad scaled area on the body.
  • Record of a spotted scat, Scatophagus argus (Perciformes: Scatophagidae) from off the Pacific coast of Oshamanbe, Hokkaido, Japan
    濵田幸穂, 三宅教平, 河合俊郎
    日本生物地理学会会報 77 109 - 113 2022/12 [Refereed][Not invited]
  • Dextral osteology of Chascanopsetta lugubris (Teleostei : Pleuronectiformes : Bothidae) from the eastern Indian Ocean
    Kazuya Kuroshima, Kota Obata, Toshio Kawai
    Bulletin of the Faculty of Fisheries Sciences, Hokkaido University 72 (2) 27 - 31 2022/12 [Not refereed][Not invited]
  • Morphological intraspecific variation of a softnose skate, Bathyraja diplotaenia (Ishiyama, 1952) (Rajiformes: Arhynchobatidae)
    Kenta Nakamura, Toshio Kawai
    Bulletin of the Faculty of Fisheries Sciences, Hokkaido University 72 (1) 9 - 16 2022/08 [Not refereed][Not invited]
  • Coloration of a small Nozawa’s prickleback, Stichaeus nozawae Jordan and Snyder, 1902 (Zoarcoidei : Stichaeidae)
    Obata, K, T. Kawai, T. Takatsu
    Bulletin of the Faculty of Fisheries Sciences, Hokkaido University 71 (1) 15 - 20 2021/08 [Not refereed][Not invited]
  • 樋口淳也, 河合俊郎, 宇治利樹, 今井圭理
    北海道大学水産科学研究彙報(Web) 71 (1) 21 - 28 2435-3353 2021/08 [Not refereed][Not invited]
  • Note on color variations of inner surface of pectoral fins in Lepidotrigla microptera Günther, 1873 (Actinopterygii: Triglidae) from Mutsu Bay, Japan
    N. Kawakami, T. Kawai, M. Nakaya
    The Thailand Natural History Museum Journal 15 (1) 51 - 58 2021/06 [Refereed][Not invited]
  • First record of Satyrichthys moluccensis (Bleeker, 1850) (Actinopterygii: Teleostei: Peristediidae) from Samoa, the Central Pacific
    J. Higuchi, T. Kawai
    The Thailand Natural History Museum Journal 14 (2) 147 - 152 2020/12 [Refereed][Not invited]
  • First record of an armored searobin, Satyrichthys clavilapis Fowler, 1938 (Teleostei: Peristediidae) from Taiwan
    T. Kawai, H.-C. Ho
    Platax 17 113 - 117 2020/12 [Refereed][Not invited]
  • TylosurinaeおよびBeloninaeに適用する標準和名の提案
    外山太一郎, 今村 央, 河合俊郎
    魚類学雑誌 67 (2) 299 - 300 2020/11 [Refereed][Not invited]
  • Deep-sea fishes from the Andaman Sea by R/V Chakratong Tongyai during 1996–2000. Part 6: orders Pleuronectiformes and Tetraodontiformes
    T. Kawai, F. Tashiro, N. Nakayama, C. Aungtonya, S. Banchongmanee
    Phuket Marine Biological Center Research Bulletin 77 101 - 108 2020/06 [Refereed][Not invited]
  • Deep-sea fishes from the Andaman Sea by R/V Chakratong Tongyai during 1996–2000. Part 5: order Perciformes
    T. Kawai, F. Tashiro, N. Nakayama, H. Imamura, K. Kamiyama, C. Aungtonya, S. Banchongmanee
    Phuket Marine Biological Center Research Bulletin 77 43 - 59 2020/06 [Refereed][Not invited]
  • Deep-sea fishes from the Andaman Sea by R/V Chakratong Tongyai during 1996–2000. Part 4: order Argentiniformes
    T. Senda, T. Kawai, F. Tashiro, H. Imamura, C. Aungtonya, S. Banchongmanee
    Phuket Marine Biological Center Research Bulletin 77 109 - 119 2020/06 [Refereed][Not invited]
  • Deep-sea fishes from the Andaman Sea by R/V Chakratong Tongyai during 1996–2000. Part 3: orders Albuliformes, Ateleopodiformes and Lampriformes
    T. Kawai, F. Tashiro, N. Nakayama, C. Aungtonya, S. Banchongmanee
    Phuket Marine Biological Center Research Bulletin 77 1 - 8 2020/06 [Refereed][Not invited]
  • Phylogenetic systematics of the needlefishes (Beloniformes: Belonidae)
    Taichirou Toyama, Toshio Kawai, Hisashi Imamura
    The Thailand Natural History Museum Journal, Monograph 1 1 - 73 2020/05 [Refereed][Not invited]
  • Coelorinchus posteromaculatus (Actinopterygii, Gadiformes, Macrouridae), a new species of grenadiers from the eastern Indian Ocean
    N. Nakayama, A. Prokofiev and T. Kawai
    Ichthyological Research 2020/03 [Refereed][Not invited]
  • カラスダラHalargyreus johnsoniiの北海道南部沖太平洋からの記録および本種の地理的変異と日本における分布域の再検討
    千田哲朗・河合俊郎・高木省吾
    日本生物地理学会会報 74 38 - 42 2019/12 [Refereed][Not invited]
  • Records of four species of armored searobins (Teleostei: Peristediidae) from Dongsha Atoll in the South China Sea
    T. Kawai and H.-C. Ho
    Platax 16 67 - 75 2019/12 [Refereed][Not invited]
  • Veera Vilasri, Hsuan-Ching Ho, Toshio Kawai, Martin F. Gomon
    Zootaxa 4702 (1) 49 - 59 2019/12 [Refereed][Not invited]
  • Katsuya Kimura, Toshio Kawai, Charatsee Aungtonya
    Phuket Marine Biological Center Research Bulletin 76 85 - 90 2019/06 [Refereed][Not invited]
  • Saki Kishimoto, Toshio Kawai, Fumihito Tashiro, Charatsee Aungtonya
    Phuket Marine Biological Center Research Bulletin 76 9 - 17 2019/06 [Refereed][Not invited]
  • Katsuya Kimura, Toshio Kawai, Fumihito Tashiro, Naohide Nakayama, Charatsee Aungtonya, Surapong Banchongmanee
    Phuket Marine Biological Center Research Bulletin 76 1 - 8 2019/06 [Refereed][Not invited]
  • Toshio Kawai
    The Thailand Natural History Museum Journal 13 (1) 69 - 75 2019/06 [Refereed][Not invited]
  • 木村克也, 今村央, 河合俊郎
    魚類学雑誌 66 (1) 121 - 122 2019/04 [Refereed][Not invited]
  • Toshio Kawai
    Ichthyological Research 2019/04 [Refereed][Not invited]
  • Tomohiro Yoshida, Toshio Kawai, Hiroyuki Motomura
    Ichthyological Research 2019 [Refereed][Not invited]
  • Saki Kishimoto, Toshio Kawai, Hisashi Imamura
    日本生物地理学会会報 73 181 - 186 2018/12 [Refereed][Not invited]
  • Takehiro Takahashi, Toshio Kawai, Hisashi Imamura
    日本生物地理学会会報 73 164 - 168 2018/12 [Refereed][Not invited]
  • Veera Vilasri, Pichai Sonchaeng, Sandra Raredon, Toshio Kawai, Tanya Chan-ard
    The Thailand Natural History Museum Journal 12 (2) 41 - 69 2018/12 [Refereed][Not invited]
  • Katsuya Kimura, Hisashi Imamura, Toshio Kawai
    Zootaxa 4536 (1) 1 - 72 2018/12 [Refereed][Not invited]
  • Martin F. Gomon, Toshio Kawai
    Raffles Bulletin of Zoology 66 624 - 651 2345-7600 2018/11 [Refereed][Not invited]
  • Satoshi Morishita, Toshio Kawai, Hiroyuki Motomura
    Ichthyological Research 1 - 10 1341-8998 2018/04/24 [Refereed][Not invited]
     
    A new species of rockfish, Sebastiscus vibrantus, is described on the basis of 24 specimens collected from Bali, Indonesia and Taiwan. Although closely resembling Sebastiscus tertius (Barsukov and Chen 1978) in overall body appearance, and having usually 19 pectoral-fin rays and black margined white blotches scattered on the lateral body surface, the new species has thicker unbranched pectoral-fin rays, a greater pectoral-fin base length, the third dorsal-fin soft ray usually longest, the anterior margin of the scaled suborbital area not reaching level the anterior margin of the orbit, and weakly developed nuchal spines.
  • Record of Histrio histrio (Lophiiformes: Antennariidae) from the Sea of Okhotsk, Yubetsu), Hokkaido, Japan
    Toshio Kawai, Naoki Tatsuta, Hajime Matsubara
    Bulletin of the Historical Museum of Urahoro 18 33 - 35 2018/03 [Not refereed][Not invited]
  • Veera Vilasri, Takuma Onodera, Toshio Kawai, Hisashi Imamura, Chikara Kawagoe
    The Thailand Natural History Museum Journal 12 (1) 29 - 34 2018 [Refereed][Not invited]
  • Katsuya Kimura, Toshio Kawai
    The Thailand Natural History Museum Journal 12 (1) 35 - 40 2018 [Refereed][Not invited]
  • 木村克也, 河合俊郎, 今村央
    日本生物地理学会会報 日本生物地理学会 72 209‐218 - 218 0067-8716 2018/01 [Refereed][Not invited]
  • Shihua Kang, Hisashi Imamura, Toshio Kawai
    Ichthyological Research 65 (1) 29 - 41 1341-8998 2018/01/01 [Refereed][Not invited]
     
    The monophyly of Polynemidae was evaluated and its sister relationship with Sciaenidae discussed, based on osteological and myological characters from 24 polynemid species in eight genera, with comparisons with acanthomorph fishes from literature and 86 species in 8 orders and 63 families examined. Polynemidae was inferred as a monophyletic group, strongly supported by 19 synapomorphies, including four unique characters (unnamed bone present on cephalic sensory canal extending from supratemporal, third actinost not supporting pectoral-fin rays, section A1 comprising lateral and medial elements, and division of obliquus inferioris present between lower postcleithrum and rod-like process on coracoid) in percoids. In addition, seven pectoral girdle characters were recognized, with the girdle possessing filament-like sensory rays, an adaptation to benthic life in muddy water. The sister relationship of Polynemidae and Sciaenidae was supported by six synapomorphies, including two rather rare (a single branchiostegal ray suspended by epihyal and posterior portions of pelvic bones on both sides interdigitated) and two unique characters (metapterygoid and quadrate interdigitated medially and anterior extension of the nasal canal).
  • 河合 俊郎, 木村 克也, 川畑 達, 神 正人
    北海道大学水産科学研究彙報 北海道大学大学院水産科学研究院 67 (3) 35 - 37 1346-1842 2017/12/28 [Not refereed][Not invited]
     
    A single male specimen of a bluntnose sixgill shark, Hexanchus griseus(Bonnaterre, 1788), was collected from the western North Pacific, off Aomori Prefecture, northern Japan in January 2015. This is the first record of this species from Aomori Prefecture, and the northern-most record of this species around Japanese waters.
  • 河合俊郎
    魚類学雑誌 日本魚類学会 64 (2) 175‐178 - 178 0021-5090 2017/11 [Refereed][Not invited]
  • R. Fricke, T. Kawai, T. Yato, H. Motomura
    Journal of the Ocean Science Foundation 28 90 - 102 2017 [Refereed][Not invited]
  • Katsuya Kimura, Megumi Takagishi, Toshio Kawai, Hisashi Imamura, Hsuan-Ching Ho, Taketeru Tomita, Fumiya Tanaka, Gento Shinohara
    Platax 14 46 - 54 2017 [Refereed][Not invited]
  • Toshio Kawai, Fumihito Tashiro, Hisashi Imamura, Charatsee Aungtonya
    Phuket Marine Biological Center Research Bulletin 74 23 - 32 2017 [Refereed][Not invited]
  • Toshio Kawai
    ZOOTAXA 4171 (2) 335 - 346 1175-5326 2016/09 [Refereed][Not invited]
     
    A new armored searobin, Peristedion richardsi sp. nov., is described on the basis of three specimens collected from the deep sea (485-600 m) off Java, Indonesia. This new species differs from the Indo-West Pacific congeners in having anterior edge of the 4th sensory pore of the rostral projection located anterior to the anterior edge of the premaxilla, perifacial rim prominent starting near anterior margin of lower jaw, spatulate rostral projection with very narrow interspace between both rostral projections, rostral projection width 1.33-1.73 in interspace between both rostral projections, and rostral projection with straight margin on medial side at base. It is clear that Peristedion nierstraszi Weber, 1913 is a junior synonym of Peristedion riversandersoni Alcock, 1894. In addition, a lectotype is designated for P. nierstraszi.
  • Hsuan-Ching Ho, Toshio Kawai, Kunio Amaoka
    ZOOTAXA 4121 (3) 267 - 294 1175-5326 2016/06 [Refereed][Not invited]
     
    An examination of the ceratioid anglerfishes collected on the Indian Ocean side of Indonesia during surveys in 2004-2005 have revealed 18 species in 9 genera and 6 families, including three new species: Cryptopsaras couesii (Ceratiidae); Melanocetus johnsonii (Melanocetidae); Diceratias trilobus, Bufoceratias microcephalus sp. nov., B. thele, B. shaoi, B. cf. wedli (Diceratiidae); Himantolophus danae, H. sagamius, H. nigricornis, H. macroceratoides (Himantolophidae); Oneirodes quadrinema sp. nov., O. amaokai sp. nov., O. carlsbergi, O. cristatus, Dermatias platynogaster, Chaenophryne cf. melanorhabdus (Oneirodidae); and Linophryne parini (Linophrynidae). Of these, specimens of B. shaoi, H. macroceratoides, O. cristatus and L. parini represent the second records since the species were described. A specimen of H. nigricornis represents the third record and a specimen of Dermatias platynogaster represents the fourth record. Descriptive data and notes on the geographical distribution and morphological variation are provided for each species.
  • V. Vilasri, R. Yamanaka, T. Kawai, H. Imamura, S. Ratmuangkhwang, F. Tashiro, M. Ohara
    The Thailand Natural History Museum Journal 10 117 - 123 2016 [Refereed][Not invited]
  • Hsuan-Ching Ho, Toshio Kawai, Wudianto, Fayakun Satria
    ACTA ICHTHYOLOGICA ET PISCATORIA 46 (2) 77 - 85 0137-1592 2016 [Refereed][Not invited]
     
    Background. The lophiid anglerfishes are with commercial importance, but rarely they are documented in Indonesian waters. During the Japanese and Indonesian collaborative surveys in 2004 and 2005, an anglerfish collection was made and it is the aim of this study to identify this collection, to document new records and to provide information further application usage such as fishery assessment from the study area. Materials and methods. Specimens were deposited in the HUMZ collection and examined by the authors. Counts, measurements, and photography were consistent with the commonly used methods. Results. Three genera and eight species in the family were collected, including Sladenia cf. zhui [sensu Ni, Wu et Li, 2012]; Lophiomus setigerus (Vahl, 1797); Lophiodes naresi (Gunther, 1880); Lophiodes bruchius Caruso, 1981; Lophiodes endoi Ho et Shao, 2008; Lophiodes lugubris (Alcock, 1894); Lophiodes miacanthus (Gilbert, 1905); and Lophiodes triradiatus (Lloyd, 1909). Of these, Sladenia cf. zhui, L. bruchius, L. endoi, L. lugubris, and L. miacanthus, are described in detail. Conclusion. The diversity of the lophiid anglerfish fauna is relatively low from the eastern Indian Ocean side of Indonesia. All species treated in the presently reported study represent the first records in Indonesian waters. A discussion of lophiid fauna from Indonesia is provided, along with a key to known species in Indonesia.
  • Hsuan-Ching Ho, Toshio Kawai, Fayakun Satria
    RAFFLES BULLETIN OF ZOOLOGY 63 301 - 308 0217-2445 2015/12 [Refereed][Not invited]
     
    The species of Chaunax are reported from the eastern Indian Ocean side of Indonesia based on a recent collection made in 2004 and 2005. Two new species from the genus Chaunax are described from Indonesia and the Philippines, both belonging to the Chaunax abei-species group. Chaunax gomoni sp. nov. is distinguished by its white peritoneum and the following combination of characters: large irregular green spots on dorsal surface when freshly dead, spots turning dark brown when preserved; head length 39.5-40.8% SL; 10 rakers on second gill arch; 11-14 neuromasts in pectoral series and 29-38 in lateral-line proper. Chaunax brachysomus sp. nov. is distinguished by its uniform pink body when freshly dead, mixed broad-based and narrow-based spines on its ventral surface, large head and short tail resulting a relatively stout body; 12 or 13 neuromasts in pectoral series and 33 in lateral-line proper; and, 9 rakers on the second gill arch. Three additional species are reported for the first time from Indonesia.
  • Annotated checklist of marine fishes from Phuket and Ranong, Thailand
    Vilasri, V, T. Yamanaka, S. Tochino, T. Kawai, S. Ratmuangkhwang, H. Imamura
    Tropical Natural History 15 (1) 55 - 68 2015 [Refereed][Not invited]
  • 河合 俊郎
    NSUGAF The Japanese Society of Fisheries Science 81 (4) 750 - 750 0021-5392 2015 [Not refereed][Not invited]
  • 荻本啓介, 河合俊郎, 松原創, 久原悠生, 永田隆一, 桑原尚司, 片倉靖次
    日本生物地理学会会報 日本生物地理学会 69 197 - 201 0067-8716 2014/12 [Refereed][Not invited]
  • Toshio Kawai
    ZOOTAXA 3900 (1) 135 - 140 1175-5326 2014/12 [Refereed][Not invited]
     
    Satyrichthys kikingeri Pogoreutz, Vitecek & Ahnelt, 2013 was described as a new species based on a single dried specimen from the Maldives, Indian Ocean. Although this species was considered to differ from the most similar species of the genus, S. laticeps (Schlegel, 1852), by numbers of barbels and anal fin rays, and several morphometric characters, it is clear that both species are inseparable as these characters were misinterpreted in the holotype in the original description of S. kikingeri. Therefore, S. kikingeri is a junior synonym of S. laticeps.
  • 佐々木 嘉子, 河合 俊郎, 三宅 教平
    日本生物地理学会会報 日本生物地理学会 69 193 - 196 0067-8716 2014/12 [Refereed][Not invited]
  • 櫻井 慎大, 大橋 慎平, 河合 俊郎
    日本生物地理学会会報 日本生物地理学会 69 117 - 124 0067-8716 2014/12 [Refereed][Not invited]
  • NAKATA Kazuyoshi, NAGANO Yuki, OHASHI Shinpei, KAWAI Toshio, OHTAKA Akifumi
    Japanese Journal of Benthology 日本ベントス学会 69 (2) 90 - 94 1345-112X 2014/12 [Refereed][Not invited]
     
    In Lake Akan, Hokkaido, northern Japan, the local population of the native and endangered Japanese crayfish <i>Cambaroides japonicus </i>(De Haan, 1841) is extinct, whereas the signal crayfish <i>Pacifastacus leniusculus </i>(Dana, 1852), an invasive species from North America, has become established and rapidly increased. This study provides information about specimens of <i>C. japonicus</i> preserved by Dr. Saburo Hatta from the lake in 1872. The six specimens (two males and four non-ovigerous females) are valuable evidence of <i>C. japonicus</i> formerly inhabiting Lake Akan. The body sizes of the specimens were 24.4–29.5 mm in carapace length and 55.7–67.4 mm in total length, and the estimated age of the largest specimen was ten years, indicating that Lake Akan of that time provided a suitable habitat. Twenty-two ectosymbiotic crayfish worms (Annelida, Clitellata, Branchiobdellida) were found attached to the specimens. Three branchiobdellidan species were identified: <i>Cirrodrilus cirratus</i> Pierantoni, 1905, <i>C. inukaii </i>(Yamaguchi, 1934) and <i>C. megalodentatus </i>(Yamaguchi, 1934). This is the first record of the latter two species from Lake Akan.
  • Ono Makoto, Kawai Toshio
    Species diversity 日本動物分類学会 19 (2) 117 - 131 1342-1670 2014/11 [Refereed][Not invited]
     
    The genus Peristedion Lacepede, 1801 occurring around Japanese waters is reviewed, with 4 species described based on 157 Japanese specimens: Peristedion orientale Temminck and Schlegel, 1843, P. liorhynchus (Gunther, 1872), P. nierstraszi Weber, 1913, and P. amblygenys Fowler, 1938. The four species are distinguished from each other by the shape of the rostral projection, the position of the fourth sensory pore on the rostral projection, the number of branches on the filamentous barbel on the lip, the total number of chin barbels, and the shape of the perifacial rim. The present study records P. amblygenys in Japanese waters for the first time.
  • 大橋 慎平, 永野 優季, 加藤 克, 河合 俊郎, 矢部 衞
    北海道大学水産科学研究彙報 北海道大学大学院水産科学研究科 64 (2) 55 - 69 1346-1842 2014/07/31 [Not refereed][Not invited]
     
    本稿で整理，紹介される標本群は，北海道大学農学部旧応用動物学教室(現動物生態学研究室)の標本室で管理されてきたもので，2011年に北海道大学総合博物館分館水産科学館に移管されたものである。まずは，この標本室における魚類標本コレクションの成立過程について整理し，関わった研究者などの情報について紹介することとしたい。
  • Taxonomy of mesopelagic fishes collected around the Ogasawara islands by the T/S Oshoro-maru
    N. Tatsuta, H. Imamura, K. Nakaya, T. Kawai, T. Abe, K. Sakaoka, S. Takagi, M. Yabe
    Memoirs of the faculty of fisheries sciences, Hokkaido University 56 (1) 1 - 64 2014/05 [Not refereed][Not invited]
  • T. Tomita, T. Kawai, H. Matsubara, M. Kobayashi, S. Katakura
    JOURNAL OF FISH BIOLOGY 84 (1) 243 - 246 0022-1112 2014/01 [Refereed][Not invited]
     
    The whale shark Rhincodon typus is the world's largest fish and it occurs in tropical, subtropical and warm temperate waters. Here, the northernmost record of R. typus is reported, when it was found in the Sea of Okhotsk for the first time. This occurrence can be explained by the unusually high sea surface temperature during the summer of 2012.
  • T. Tomita, T. Kawai, H. Matsubara, R. Nagata
    Journal of Fish Biology 83 (3) 695 - 698 0022-1112 2013/09 [Refereed][Not invited]
     
    The northernmost record for Chilean devil ray Mobula tarapacana, a circumglobal species that occurs in tropical, subtropical and limited warm temperate waters, is described. An adult female was caught incidentally in the Sea of Okhotsk on 17 September 2011. This specimen is the first confirmed occurrence of devil rays Mobula spp. in cold temperate waters. © 2013 The Fisheries Society of the British Isles.
  • Hisashi Imamura, Toshio Kawai
    ICHTHYOLOGICAL RESEARCH 60 (3) 291 - 292 1341-8998 2013/07 [Refereed][Not invited]
  • Martien J.P. Van Oijen, Toshio Kawai, Ineke Loots
    Zootaxa 3670 (2) 207 - 214 1175-5326 2013/06/12 [Refereed][Not invited]
     
    The presumed type status of the two remaining specimens of eristedion moluccense Bleeker 1850 in the collection the Naturalis Biodiversity Center was examined by comparing them to descriptions in two Bleeker papers and an unpublished manuscript by Bleeker written for the Atlas Ichthyologique. Latin, French and Dutch parts of these papers were translated into English. The Dutch description of P. laticeps Schlegel 1852 was also translated. In the course of this study it became clear why Bleeker considered P. laticeps, a as a synonym of P. moluccense. Copyright © 2013 Magnolia Press.
  • Toshio Kawai
    ZOOTAXA 3635 (4) 419 - 438 1175-5326 2013/03 [Refereed][Not invited]
     
    The Indo-Pacific peristediid genus Satyrichthys Kaup, 1873 was first diagnosed as having a broad head with mesethmoid, postocular, parietal and preopercular spines. Later, most ichthyologists characterized Satyrichthys by its toothless jaws and strong preopercular spine. Kawai (2008) divided Satyrichthys into two genera, Satyrichthys and Scalicus, on the basis of a phylogenetic hypothesis, and redefined Satyrichthys. Seven species of Satyrichthys are recognized here, including one new species: S. clavilapis, S. laticeps, S. longiceps, S. milleri sp. nov., S. moluccensis, S. rieffeli and S. welchi. The new species is distinguished from its congeners in having equilateral-triangular rostral projections, 4 lip and 4 chin barbels, and no anterior directed spines on the upper lateral row of the caudal peduncle. Satyrichthys laticeps, previously treated as a junior synonym of S. moluccensis, is regarded as a valid species. The following new synonyms are noted: S. adeni, S. halyi and S. magnus are all junior synonyms of S. laticeps; S. isokawae is a junior synonym of S. moluccensis; and S. lingi is a junior synonym of S. welchi. In addition, a neotype is designated for S. moluccensis.
  • Yasuhiro Honma, Hisashi Imamura, Toshio Kawai
    Ichthyological Research 60 (2) 122 - 141 1341-8998 2013 [Refereed][Not invited]
     
    Osteological and myological characters of the genus Perryena are described and the phylogenetic position of the genus is inferred cladistically on the basis of 89 character transformation series, with Perryena representing a branch off a clade comprising the Tetrarogidae, Synanceiidae, Aploactinidae, Congiopodidae, Gnathanacanthidae and Pataecidae, following an initial divergence of the family Apistidae. Perryena is not closely related to Congiopodidae, despite its historical referral to that family. In keeping with the familial rankings given to successive branches of the remaining ingroup, the new family name Perryenidae is proposed for Perryena, here designated as the type genus of the family. © 2013 The Ichthyological Society of Japan.
  • 櫻井 慎大, 佐々木 嘉子, 河合 俊郎
    日本生物地理学会会報 日本生物地理学会 67 241 - 246 0067-8716 2012/12 [Refereed][Not invited]
  • TENDA Miki, KAWAI Toshio
    Species Diversity 日本動物分類学会 17 (2) 135 - 144 1342-1670 2012/11 [Refereed][Not invited]
     
    A large series of Peristedion barbiger (Garman, 1899) was collected in the eastern Pacific off Peru during a joint survey conducted by the Japan Deep Sea Trawlers Association and the Instituto del Mar del Peru in 1998-2003. No detailed description of P. barbiger has been published since the original description, and the availability of 285 new specimens provided an opportunity to determine the extent morphological variability within this species. Our study revealed that the presence or absence of nasal spines and the shape of the preopercular margin, both having been used previously as taxonomic characters, vary intraspecifically. The species is diagnosed as having 16-19 dorsal-fin soft rays; 31-34 bony plates in the upper lateral row; 14-19 lower gill rakers; 17-25 chin barbels in total; short pectoral fin, 13.3-17.8% of SL; a relatively short pair of filamentous barbel on the lip, 26.4-57.1% of HL; a broad, spatulate rostral projection of a length 19.7-39.5% of HL; and a large black spot on the spinous portion of the dorsal fin. In addition, we herein designate a lectotype for P. barbiger.
  • Matsubara Hajime, Kawai Toshio, Iwata Daisuke
    Biogeography Biogeographical Society of Japan 14 11 - 17 1345-0662 2012/09 [Refereed][Not invited]
  • Toshio Kawai, Kunio Amaoka, Bernard Seret
    ZOOTAXA (3135) 63 - 68 1175-5326 2011/12 [Refereed][Not invited]
     
    A new righteye flounder, Samariscus neocaledonia sp. nov., is described on the basis of two specimens collected in deep waters (244-278 m) around New Caledonia. The new species is easily distinguished from its 18 congeners in having a combination of 78-81 dorsal fin rays, 62-65 anal fin rays, five pectoral fin rays, ca. 55-62 lateral line scales, and 10 abdominal and 31-32 caudal vertebrae.
  • 河合俊郎, 松原創
    北海道大学水産科学研究い報 北海道大学大学院水産科学研究院 = Research Faculty of Fisheries Sciences, Hokkaido University 61 (1) 1 - 4 1346-1842 2011/06/20 [Not refereed][Not invited]
     
    A single specimen of a hermaphroditic tiger puffer, Takifugu rubripes (Temminck and Schlegel, 1850), was collected from the western North Pacific, off Hokuto, Hokkaido, Japan. The left gonad of the specimen was of testicular tissue, and the right gonad was ovarian tissue sandwiched by testicular tissue. This is the first instance of hermaphrodism in this species. In addition, the anterior and posterior nostrils of the specimen were fused to each other, and the left side of the body was abnormally pigmented.
  • 長尾太地, 河合俊郎, 松原創, 鈴木淳志
    知床博物館研究報告 (32) 31 - 34 0387-8716 2011/03/31 [Not refereed][Not invited]
  • Toshio Kawai
    ICHTHYOLOGICAL RESEARCH Springer Japan 58 (1) 67 - 71 1341-8998 2011/01 [Refereed][Not invited]
     
    Peristedium indicum Brauer 1906 (family Peristediidae) has not been reported since its original description. Reexamination of syntypes of P. indicum revealed that P. indicum is included in the genus Paraheminodus Kamohara 1957 because of having upper jaw teeth, a smooth lateral margin, posterior parts of lower lateral rows of bony plates separated from each other, and all lower jaw barbels unbranched except for the posteriormost barbels. Based on a comparison of syntypes of P. indicum with all types and other materials of all congeners of Paraheminodus, it is concluded that P. indicum is a junior synonym of Paraheminodus murrayi (Gunther 1880).
  • NAGANO Yuki, TADA Tomoyo, TANI Takashi, KAWAI Toshio
    Bulletin of the Biogeographical Society of Japan 日本生物地理学会 65 51 - 56 0067-8716 2010/12 [Refereed][Not invited]
  • Toshio Kawai, Kunio Amaoka, Bernard Seret
    ICHTHYOLOGICAL RESEARCH 57 (2) 193 - 198 1341-8998 2010/04 [Refereed][Not invited]
     
    A new righteye flounder, Poecilopsetta multiradiata, is described from eight specimens (two males and six females) collected from deep waters (336-408 m) around New Zealand and New Caledonia (South-West Pacific). This new species is distinguished from its 14 congeners by the following combination of characters: high numbers of dorsal (70-73) and anal (58-62) fin rays, ca. 85-99 lateral-line scales, 31-32 caudal vertebrae, and a relatively shallow body depth of 36.9-41.9% SL.
  • 高田陽子, 河合俊郎, 松原創, 鈴木淳志
    日本生物地理学会会報 日本生物地理学会 64 13 - 19 0067-8716 2009/12/20 [Not refereed][Not invited]
  • KAWAI Toshio, TAKATA Yohko
    Species Diversity 日本動物分類学会 14 (2) 89 - 96 1342-1670 2009/07 [Refereed][Not invited]
     
    Satyrichthys rugosus (Fowler, 1943) was established with no comparison to other peristediid searobins. Our examination of types and other material of both S. rugosus and S. clavilapis (Fowler, 1938) has clarified that S. rugosus represents postlarvae and juveniles of the latter species. We therefore regard S. rugosus as a junior synonym of S. clavilapis and comment on its ontogenetic changes in morphology.
  • 河合俊郎, 佐藤崇, 松原創, 鈴木淳志
    日本生物地理学会会報 63 47 - 50 0067-8716 2008/12 [Refereed][Not invited]
  • Kawai Toshio, Shinohara Gento
    Bulletin of the National Museum of Nature and Science. Series A, Zoology 国立科学博物館 34 (4) 175 - 181 1881-9052 2008/12 [Refereed][Not invited]
     
    A rare carapid fish, Eurypleuron owasianum, and a rare chiasmodontid fish, Dysalotus alcocki, were collected from the Okinawa Trough, East China Sea in 2002 and 2003, respectively. Adult specimens of E. owasianum have only been recorded in Japan from its type locality, Owase, Mie Prefecture. No specimens of D. alcocki have been reported from Japanese waters, although this species has been included in Japanese checklists. We report morphological characters of both specimens and include color photographs.
  • Toshio Kawai, Kazuhiro Nakaya, Bernard Seret
    ICHTHYOLOGICAL RESEARCH 55 (4) 374 - 378 1341-8998 2008/11 [Refereed][Not invited]
     
    A new species of armored searobin, Paraheminodus longirostralis, is described from five specimens collected from New Caledonia at depths of 412-467 m. It is distinguishable from its three known congeners in having 34 bony plates in the upper lateral row, a forward-directed spine on each plate between the 23rd-26th and 31st-32nd plates in the upper lateral row, 6-7 + 1 + 20-21 = 27-28 gill rakers, an elongate body posterior to the anus (49.9-52.1% standard length), an elongate rostral projection (53.0-59.3% head length), short upper jaw (42.1-43.4% head length), an elongate pectoral fin (70.6-79.4% head length), and long preopercular spine (39.2-57.7% head length).
  • KAWAI Toshio, TAKATA Yohko
    Biogeography Biogeographical Society of Japan 10 27 - 31 1345-0662 2008/08 [Refereed][Not invited]
  • Kawai Toshio, Tashiro Fumihito
    Jpn. J. Ichthyol. The Ichthyological Society of Japan 55 (1) 43 - 47 1884-7374 2008/05 [Refereed][Not invited]
     
    A single peristediid specimen, NSMT-P 79556, was caught from Suruga Bay, Japan, and identified as <I>Satyrichthys adeni</I> (Lloyd, 1907) based on the following combination of characters: 16 dorsal and anal fin soft rays, respectively; 4 barbels on lip and 3 on chin; rostral projection length 9.9% SL; no straight sutured line formed by both frontals and parietals on posterior dorsal head; no black spots on dorsal fin; and pectoral fin pale yellow in alcohol. The present study represents the first record of this species in Japanese waters.
  • KAWAI Toshio, YATO Takuji
    魚類学雜誌 55 (1) 61 - 62 0021-5090 2008/05 [Refereed][Not invited]
  • KAWAI Toshio
    Species Diversity 日本動物分類学会 13 (1) 1 - 34 1342-1670 2008/05 [Refereed][Not invited]
     
    The family Peristediidae, comprising about 36 species of armored sea robins in five genera, inhabit the bottoms of the tropical and temperate waters of the world oceans in depths of about 50 to 800 m. The aim of this study is to infer relationships among the species in the family based on morphological characters and to revise the genus-level classification on the basis of the inferred pattern of phylogeny. Twenty-four peristediid and 16 outgroup taxa, i.e., 13 triglid and three hoplichthyid species, were used for the phylogenetic analysis. Monophyly of the Peristediidae is highly corroborated and two major clades are recognized in the family. The first clade includes Gargariscus, Heminodus, Satyrichthys, and Paraheminodus, and the second includes only Peristedion. It became clear that the genus Satyrichthys is a nonmonophyletic group. In conclusion, six monophyletic genera are recognized in the Peristediidae: Gargariscus, Heminodus, Paraheminodus, Peristedion, Satyrichthys, and Scalicus. Accordingly, I propose seven new combinations with Scalicus as follows: Scalicus engyceros, S. gilberti, S. hians, S. investigatoris, S. orientalis, S. quadratorostratus, and S. serrulatus.
  • Toshio Kawai, Kunio Amaoka, Bernard Seret
    ICHTHYOLOGICAL RESEARCH 55 (1) 17 - 21 1341-8998 2008/02 [Refereed][Not invited]
     
    A new dextral flounder, Samariscus multiradiatus, is described from six specimens (four males and two females) collected in deep waters (296-430 m) around New Caledonia. The species is easily distinguished from its 16 congeners in having a combination of 85-91 dorsal fin rays, 67-72 anal fin rays, 5 pectoral fin rays, and 9 abdominal and 34-35 caudal vertebrae.
  • KAWAI Toshio, NAKAYA Kazuhiro
    Species Diversity 日本動物分類学会 12 (3) 167 - 173 1342-1670 2007/09 [Refereed][Not invited]
     
    A rare peristediid fish, Heminodus philippinus Smith, 1917, is redescribed on the basis of the holotype and 25 additional specimens collected from the eastern Indian Ocean off Java and Sumatra in Indonesia. Our examination has revealed that H. japonicus Kamohara, 1952, described from Japan, is a junior synonym of H. philippinus.
  • Kunio Amaoka, Toshio Kawai, Bernard Seret
    ICHTHYOLOGICAL RESEARCH 53 (3) 223 - 227 1341-8998 2006/08 [Refereed][Not invited]
     
    A new poecilopsettid flounder, Nematops nanosquama, is described from 10 specimens (4 males, 6 females) collected from deep waters (96-650m) off Hiva Oa, Marquesas Islands. This species is easily separated from the three recognized species of the genus Nematops by having large numbers of dorsal fin rays, anal fin rays, lateral line scales, and vertebrae, five dark transverse broad bands on the body, and a black blotch on the distal area of the pectoral fin. N.nanosquama shows the easternmost record of this genus from the Pacific Ocean.
  • Toshio Kawai, Kunio Amaoka
    ICHTHYOLOGICAL RESEARCH 53 (3) 264 - 268 1341-8998 2006/08 [Refereed][Not invited]
     
    A new righteye flounder, Poecilopsetta pectoralis, is described from 4 specimens (2 males, 2 females) collected from deep waters (510-580m) around New Caledonia. The species is easily distinguished from 13 congeners by having 99-105 lateral line scales, a narrow interorbit [male: 37.00-40.25 in head length (HL); female: 47.33-50.71], long pectoral fin on ocular side (male: 0.89-0.91 in HL; female: 1.04-1.20), no scales from snout to anterior interorbital region, and 4 spots on caudal fin. Sexual dimorphism is evident in this species, showing males bearing the ocular-side pectoral fin longer than females, and it is the first example in Poecilopsetta.
  • KAWAI Toshio, YABE Mamoru
    Species Diversity 日本動物分類学会 11 (1) 1 - 5 1342-1670 2006/02 [Refereed][Not invited]
     
    Peristedion picturatum McCulloch, 1926 (family Peristediidae) was distinguished from P. liorhynchus (Gunther, 1872) by a single character, the presence of two bony plates between the anus and first anal ray (vs. three in P. liorhynchus). Our examination of the holotype of P. liorhynchus revealed that it also actually has just two bony plates there, not three. Based on a comparison of the holotype and other material of P. picturatum with the holotype of P. liorhynchus, we conclude that P. picturatum is a junior synonym of P. liorhynchus.
  • KAWAI Toshio, TAKAYA Fumihito, YABE Mamoru
    Bulletin of the Biogeographical Society of Japan 日本生物地理学会 59 (0) 11 - 13 0067-8716 2004/12 [Refereed][Not invited]
  • T Kawai, H Imamura, K Nakaya
    ICHTHYOLOGICAL RESEARCH 51 (2) 126 - 130 1341-8998 2004/05 [Refereed][Not invited]
     
    A new peristediid, Paraheminodus kamoharai, is described from three specimens (106.9-114.8 mm SL) collected from the Sulu Sea, the Philippines. The new species is easily distinguished from its congeners (Paraheminodus laticephalus and P. murrayi) in having 33 bony plates in the upper lateral row, the 24-25th to 30th bony plate each in the upper lateral row with a forwardly directed spine, 17-19 gill rakers on the lower limb, a long slender rostral projection (43.4-47.0% of head length), and short upper (41.1-42.3%) and lower jaws (36.5-37.2%).
  • T Kawai, H Imamura, K Nakaya
    ICHTHYOLOGICAL RESEARCH 51 (1) 73 - 76 1341-8998 2004/02 [Refereed][Not invited]
     
    Paraheminodus kochiensis Kamohara, 1957 (family Peristediidae), has been regarded by most workers as a junior synonym of P. murrayi (GUnther, 1880), but sufficient argument for combining the two species has not been made. A detailed comparison of the type and non-type material of P. kochiensis and P. murrayi revealed no significant differences between the two; thus, their synonymy is hereby confirmed. Paraheminodus murrayi and P. laticephalus (Kamohara, 1952) are clearly distinguished by the morphology of the upper jaw teeth, length of the upper jaw, and by the number of barbels.
  • Kawai Toshio, Imamura Hisashi, Nakaya Kazuhiro
    Bulletin of Fisheries Sciences, Hokkaido University 北海道大学 54 (1) 13 - 16 1346-1842 2003/07 [Not refereed][Not invited]
  • Toshio Kawai, Hisashi Imamura, Yoshio Ishito, Kazuhiro Imamura
    北海道大学水産科学研究彙報 北海道大学 53 (3) 83 - 86 1346-1842 2002/12 [Not refereed][Not invited]
     
    2002年に青森県八戸市沖から各1個体のアナゴ科魚類およびキンメダイ科魚類が採集された。これらの標本を分類学的に詳細に調べた結果、ゴテンアナゴariosoma meekiおよびナンヨウキンメBeryx decadactylus Cuvier、1829と査定された。ゴテンアナゴは太平洋およびインド洋に分布し、日本では南日本各地、新潟県、岩手県などから報告されている。。ナンヨウキンメは世界の熱帯から温帯にかけての深海に分布する。本種は多くの著者によって南日本から報告されており、北日本では新潟県および岩手県からの2報告例しか知られていない。本研究では八戸市沖で採集されたゴテンアナゴおよびナンヨウキンメを青森県からの初記録種とするとともに、これらの種の北限記録として報告する。

Books etc

  • 北水ブックス 道南おさかな図鑑
    魚に魅せられた北大の研究者たち (Joint workカナガシラ、ハダカイワシ類)
    海文堂出版 2023/10 (ISBN: 9784303800116)
  • Coastal fishes of the western Indian Ocean
    Toshio Kawai, William J. RIchards (ContributorPeristediidae)
    South African Institute for Aquatic Biodiversity 2022/09
  • 水産科学・海洋環境科学実習
    河合俊郎 (Contributor1.1噴火湾沖で採集される深海性魚類)
    海文堂出版 2019/07 (ISBN: 9784303115005)
  • 魚類学の百科事典
    河合俊郎 (Contributor魚類の多様性、条鰭類の鰭)
    丸善出版 2018/10
  • 魚類学
    河合俊郎 (Contributor3章 体表の構造)
    恒星社厚生閣 2017/09
  • Commercial and bycatch market fishes of Ponay Island, Philippines
    Toshio Kawai (ContributorTriglidae, Peristediidae)
    Kagoshima University Museum 2017/01
  • 生物の形や能力を利用する学問 バイオミメティクス
    篠原現人, 松浦啓一, 河合俊郎 (Contributor魚類のかたちと生息環境)
    東海大学出版部 2016/03
  • 学船 北海道大学 洋上のキャンパスおしょろ丸
    河合俊郎, 藤田良治, 中原隆史 (Contributorおしょろ丸の概要と歴史)
    中西出版 2014/07
  • 次世代バイオミメティクス研究の最前線―生物多様性に学ぶ―
    篠原現人, 松浦啓一, 河合俊郎 (Contributor第5章 2バイオミメティック・データベースとしての魚類インベントリー)
    シーエムシー出版 2011/08
  • 新魚類解剖図鑑
    河合俊郎 (Contributor循環器系・内臓)
    緑書房 2010
  • ペルー海域の深海魚類図鑑
    河合俊郎 (Contributorホウボウ科・キホウボウ科)
    （社）日本トロール底魚協会 2009
  • 北海道大学大学院文学研究科芸術学研究室, 北海道大学大学院文学研究科芸術学講座, 北海道大学総合博物館, 疋田 豊治, 廣田 理紗, 疋田 豊彦, 港 千尋, 大下 智一, 河合 俊郎, 北村 清彦 
    北海道大学大学院文学研究科芸術学研究室 : 北海道大学総合博物館 2009

Conference Activities & Talks

Works

Research Grants & Projects

  • 日本学術振興会：科学研究費助成事業
    Date (from‐to) : 2023/04 -2027/03 
    Author : 河合 俊郎
  • Japan Society for the Promotion of Science：Grants-in-Aid for Scientific Research
    Date (from‐to) : 2012/06 -2017/03 
    Author : NOMURA SHUHEI, OGAWA TAKAHIRO, TSUCHIYA HIROSHI, KAWAI TOSHIO
     
    Biologists of this team accumulated more than 30,000 datasets, composed of SEM photos and text data on insects, birds or fishes. Information scientists built a biomimetic image retrieval system supported by ontology. This system is opened to the public through the internet. Outreach activities including an exhibition on biomimetics in the National Museum of Nature and Science, Tokyo (April-June 2016) were carried out. A book on biomimetics for general public was published in March 2016.

Educational Activities

Teaching Experience

  • Oggs PARE program summer courceOggs PARE program summer cource Hokkaido University
  • Oggs PARE program spring courceOggs PARE program spring cource Hokkaido University
  • Oggs PARE program summer courceOggs PARE program summer cource Hokkaido University
  • Aquatic Biology
    開講年度 : 2021
    課程区分 : 学士課程
    開講学部 : 水産学部
    キーワード : 学名、命名法、系統樹、分類体系、学術標本、種多様性
  • Field Training on Aquatic Biological Science
    開講年度 : 2021
    課程区分 : 学士課程
    開講学部 : 水産学部
    キーワード : 生物相・分類・生態・物質循環・行動・組織
  • Shipboard Training
    開講年度 : 2021
    課程区分 : 学士課程
    開講学部 : 水産学部
    キーワード : おしょろ丸、海洋観測、生物採集、目視観察
  • Marine Science Biology II - Laboratory
    開講年度 : 2021
    課程区分 : 学士課程
    開講学部 : 水産学部
    キーワード : サケ　マス　ベントス　貝類　甲殻類　魚類　頭足類 分類　成長曲線　資源量推定　生物測定　外部形態　稚魚　耳石
  • Marine Science Biology III - Laboratory
    開講年度 : 2021
    課程区分 : 学士課程
    開講学部 : 水産学部
    キーワード : 魚類、プランクトン、解剖、分類、形態、生態、生?、サケ
  • Teaching Method of School Subjects(Fishery Ⅰ)
    開講年度 : 2021
    課程区分 : 学士課程
    開講学部 : 教育学部

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