研究者データベース

李 相逸(イ サンイル)
工学研究院 環境工学部門 環境工学
助教

基本情報

所属

  • 工学研究院 環境工学部門 環境工学

職名

  • 助教

学位

  • 博士(感性学)(九州大学)

ホームページURL

J-Global ID

研究キーワード

  • 住環境   光環境   時間生物学   生理人類学   環境人間工学   

研究分野

  • ライフサイエンス / 栄養学、健康科学
  • 人文・社会 / 家政学、生活科学
  • ライフサイエンス / 応用人類学

職歴

  • 2017年12月 - 現在 北海道大学 大学院工学研究院 助教
  • 2016年04月 - 2017年11月 九州大学 大学院芸術工学研究院 特任助教
  • 2014年04月 - 2016年03月 九州大学 大学院芸術工学研究院 学術研究員

所属学協会

  • 空気調和・衛生工学会   日本睡眠学会   日本時間生物学会   日本生理人類学会   日本建築学会   

研究活動情報

論文

  • Melatonin suppression during a simulated night shift in medium intensity light is increased by 10-minute breaks in dim light and decreased by 10-minute breaks in bright light
    Sang-Il Lee, Saki Kinoshita, Anna Noguchi, Taisuke Eto, Michihiro Ohashi, Yuki Nishimura, Kaho Maeda, Yuki Motomura, Yasuhiro Awata, Shigekazu Higuchi
    Chronobiology International 2020年04月 [査読有り][通常論文]
  • Melatonin suppression and sleepiness in children exposed to blue-enriched white LED lighting at night
    Sang-il Lee, Kouhei Matsumori, Kana Nishimura, Yuki Nishimura, Yuki Ikeda, Taisuke Eto, Shigekazu Higuchi
    Physiological Reports 6 24 e13942  2018年12月 [査読有り][通常論文]
  • 瞳孔の対光反射のスペクトル感度に関する研究:児童と若年成人の比較
    江藤太亮, 松森孝平, 李相逸, 樋口重和
    日本生理人類学会誌 23 2 63 - 67 2018年03月 [査読有り][通常論文]
  • L-Serine Enhances Light-Induced Circadian Phase Resetting in Mice and Humans
    Shinobu Yasuo, Ayaka Iwamoto, Sang-il Lee, Shotaro Ochiai, Rina Hitachi, Satomi Shibata, Nobuo Uotsu, Chie Tarumizu, Sayuri Matsuoka, Mitsuhiro Furuse, Shigekazu Higuchi
    The Journal of Nutrition 147 12 2347 - 2355 2017年10月 [査読有り][通常論文]
  • Tokiho Akiyama, Takafumi Katsumura, Shigeki Nakagome, Sang-il Lee, Keiichiro Joh, Hidenobu Soejima, Kazuma Fujimoto, Ryosuke Kimura, Hajime Ishida, Tsunehiko Hanihara, Akira Yasukouchi, Yoko Satta, Shigekazu Higuchi, Hiroki Oota
    PLOS ONE 12 6 (e0178373)  2017年06月 [査読有り][通常論文]
     
    Humans show various responses to the environmental stimulus in individual levels as "physiological variations." However, it has been unclear if these are caused by genetic variations. In this study, we examined the association between the physiological variation of response to light-stimulus and genetic polymorphisms. We collected physiological data from 43 subjects, including light-induced melatonin suppression, and performed haplotype analyses on the clock genes, PER2 and PER3, exhibiting geographical differentiation of allele frequencies. Among the haplotypes of PER3, no significant difference in light sensitivity was found. However, three common haplotypes of PER2 accounted for more than 96% of the chromosomes in subjects, and 1 of those 3 had a significantly low-sensitive response to light-stimulus (P < 0.05). The homozygote of the low-sensitive PER2 haplotype showed significantly lower percentages of melatonin suppression (P 0.05), and the heterozygotes of the haplotypes varied their ratios, indicating that the physiological variation for light-sensitivity is evidently related to the PER2 polymorphism. Compared with global haplotype frequencies, the haplotype with a low-sensitive response was more frequent in Africans than in non-Africans, and came to the root in the phylogenetic tree, suggesting that the low light-sensitive haplotype is the ancestral type, whereas the other haplotypes with high sensitivity to light are the derived types. Hence, we speculate that the high light-sensitive haplotypes have spread throughout the world after the Out-of-Africa migration of modern humans.
  • 落合将太郎, 原田和樹, 李相逸, 樋口重和
    日本生理人類学会誌 22 2 69 - 76 2017年05月 [査読有り][通常論文]
  • Shigekazu Higuchi, Sang-il Lee, Tomoaki Kozaki, Tetsuo Harada, Ikuo Tanaka
    CHRONOBIOLOGY INTERNATIONAL 33 4 448 - 452 2016年 [査読有り][通常論文]
     
    Light is the strongest synchronizer of human circadian rhythms, and exposure to residential light at night reportedly causes a delay of circadian rhythms. The present study was conducted to investigate the association between color temperature of light at home and circadian phase of salivary melatonin in adults and children. Twenty healthy children ( mean age: 9.7 year) and 17 of their parents ( mean age: 41.9 years) participated in the experiment. Circadian phase assessments were made with dim light melatonin onset ( DLMO). There were large individual variations in DLMO both in adults and children. The average DLMO in adults and in children were 21: 50 +/- 1: 12 and 20: 55 +/- 0: 44, respectively. The average illuminance and color temperature of light at eye level were 139.6 +/- 82.7 lx and 3862.0 +/- 965.6 K, respectively. There were significant correlations between color temperature of light and DLMO in adults ( r = 0.735, p < 0.01) and children ( r = 0.479, p < 0.05), although no significant correlations were found between illuminance level and DLMO. The results suggest that high color temperature light at home might be a cause of the delay of circadian phase in adults and children.
  • 光のサーカディアンリズムとメラトニン分泌への作用の個人差
    樋口重和, 李相逸
    照明学会誌 99 1 20 - 24 2015年01月 [査読有り][招待有り]
  • Shigekazu Higuchi, Yuki Nagafuchi, Sang-il Lee, Tetsuo Harada
    JOURNAL OF CLINICAL ENDOCRINOLOGY & METABOLISM 99 9 3298 - 3303 2014年09月 [査読有り][通常論文]
     
    Context: The sensitivity of melatonin to light suppression is expected to be higher in children because children have large pupils and pure crystal lenses. However, melatonin suppression by light in children remains unclear. Objective: We investigated whether light-induced melatonin suppression in children is larger than that in adults. Methods: Thirty-three healthy primary school children (meanage, 9.2 +/- 1.5 y) and 29 healthy adults (meanage, 41.6 +/- 4.7 y) participated in two experiments. In the first experiment, salivary melatonin concentrations in 13 children and 13 adults were measured at night under a dim light (<30 lux) and a moderately bright light (580 lux) in an experimental facility. Pupil diameters were also measured under dim light and bright light. In the second experiment, melatonin concentrations in 20 children and 16 adults were measured under dim light in the experimental facility and under room light at home (illuminance, 140.0 +/- 82.7 lux). Results: In experiment 1, the melatonin concentration was significantly decreased by exposure to moderately bright light in both adults and children. Melatonin suppression was significantly larger in children (88.2%; n = 5) than in adults (46.3%; n = 6; P < .01), although the data for some participants were excluded because melatonin concentrations had not yet risen. In experiment 2, melatonin secretion was significantly suppressed by room light at home in children (n = 15; P < .05) but not in adults (n = 11). Conclusion: We found that the percentage of melatonin suppression by light in children was almost twice that in adults, suggesting that melatonin is more sensitive to light in children than in adults at night.
  • Kenta Hayashida, Shigekazu Higuchi, Jiro Kajiwara, Sang-il Lee, Kenji Kai, Katsuyoshi Setsu, Fumitada Hattori
    SLEEP AND BIOLOGICAL RHYTHMS 12 3 220 - 223 2014年07月 [査読有り][通常論文]
     
    The purpose of the present study was to clarify the relationship between nighttime sleep and effectiveness of inpatient rehabilitation for stroke patients during convalescence. Fifteen patients participated. Nighttime sleep was measured by actigraph recording. Actual sleep time was significantly correlated with discharge motor Functional Independence Measure (FIM) score and FIM efficiency. Wake after sleep onset (WASO) time was significantly correlated with length of stay. Sleep efficiency was significantly correlated with discharge motor FIM score. Our study showed a relationship between nighttime sleep and effectiveness of inpatient rehabilitation during convalescence after stroke.
  • Sang-il Lee, Akiko Hida, Shingo Kitamura, Kazuo Mishima, Shigekazu Higuchi
    JOURNAL OF PHYSIOLOGICAL ANTHROPOLOGY 33 9 2014年05月 [査読有り][通常論文]
     
    Background: In our previous studies, we found that the Ile394Thr SNP in the melanopsin gene (OPN4) was functionally associated with the pupillary light reflex. This indicates the possibility that OPN4*Ile394Thr is associated with other non-image forming responses. The aim of this study was therefore to determine whether OPN4*Ile394Thr is associated with sleep/wake timing. Methods: A total of 348 healthy Japanese university students participated in this study. Scalp hair was used to genotype the Ile394Thr SNP of OPN4. Sleep habits, including bedtime, wake time and sleep duration, were assessed separately for weekdays and weekends. A total of 328 samples, including 223 samples with TT genotype, 91 with TC genotype and 14 with CC genotype, were used for statistical analysis. No significant difference in age or male/female distribution was found among the three genotype groups. Results: There was no significant difference in circadian preference among the genotype groups. During weekdays, bedtime, wake time and midpoint of sleep for CC subjects were significantly later than those for TT and TC subjects. However, there was no difference between TT and TC subjects in any of their sleep habits. During weekends, bedtime of CC subjects was significantly later than those of TT and TC subjects, and the midpoint of sleep of CC subjects was significantly later than that of TC subjects. Conclusions: Our findings demonstrated that OPN4*Ile394Thr is associated with sleep/wake timing. We also found that the sleep/wake timing of subjects with the CC genotype was later than that of subjects with the TT or TC genotype.
  • A missense I394T Variant in Human Melanopsin Gene is Associated with Non-Image Forming Responses、including Pupillary Light Reflex and Sleep/Wake Timing
    Sang-il Lee, Shigekazu Higuchi
    The 1st Asian Conference on Ergonomics and Design (ACED) 2014年 [査読有り][通常論文]
  • Sang-il Lee, Akiko Hida, Sei-ichi Tsujimura, Takeshi Morita, Kazuo Mishima, Shigekazu Higuchi
    JOURNAL OF PHYSIOLOGICAL ANTHROPOLOGY 32 16 2013年10月 [査読有り][通常論文]
     
    Background: Our aim was to determine the association between melanopsin gene polymorphism and pupillary light reflex under diverse photic conditions, including different intensities and wavelengths. Methods: A total of 195 visually corrected subjects volunteered for investigation of the melanopsin gene of single nucleotide polymorphism (SNP) of rs1079610 (I394T). The genotype groups were TT (n = 126), TC (n = 55), and CC (n = 8), and 75 of the subjects, including subjects with TT (n = 34), TC (n = 33), and CC (n = 8) participated in our experiment. Three monochromatic lights with peak wavelengths of 465 nm (blue), 536 nm (green), and 632 nm (red) were prepared, and each light was projected to the subjects with five intensities, 12, 13, 14, 14.5 and 15 log photons/(cm2 s), for one minute. The pupil size of the left eye was measured under each light condition after a 1- minute adaptation. Results: The pupils of the TC + CC genotypes (n = 38) were significantly smaller than those of the TT genotype (n = 31) under a blue (463 nm) light condition with 15 log photons/(cm2 s) (P < 0.05). In contrast, there were no significant differences under green (536 nm) and red (632 nm) light conditions. Conversely, relative pupil constrictions of the TC + CC genotypes were greater than those of the TT genotype under both blue and green conditions with high intensities (14.5 and 15 log photons/(cm2 s)). In contrast, there were no significant differences between genotype groups in pupil size and relative pupilloconstriction under the red light conditions. Conclusions: Our findings suggest that the melanopsin gene polymorphism (I394T) functionally interacts with pupillary light reflex, depending on light intensity and, particularly, wavelength, and that under a light condition fulfilling both high intensity and short wavelength, the pupillary light response of subjects with the C allele (TC + CC) is more sensitive to light than that of subjects with the TT genotype.
  • Shigekazu Higuchi, Akiko Hida, Sei-ichi Tsujimura, Kazuo Mishima, Akira Yasukouchi, Sang-il Lee, Youhei Kinjyo, Manabu Miyahira
    PloS one 8 3 e60310  2013年 [査読有り][通常論文]
     
    BACKGROUND: Melanopsin-containing intrinsically photosensitive retinal ganglion cells (ipRGCs) play an important role in non-image forming responses to light, such as circadian photoentrainment, light-induced melatonin suppression, and pupillary light response. Although it is known that there are some single nucleotide polymorphisms (SNPs) in the melanopsin (OPN4) gene in humans, the associations of the SNPs with non-image forming responses to light remains unclear. In the present study, we examined the associations of melanopsin gene polymorphisms with pupillary light response. METHODS: Japanese university students (mean age: 21.0 ± 1.7 years) with the genotypes of TT (n = 38), TC (n = 28) and CC (n = 7) at rs1079610 (I394T) located in the coding region participated in the present study. They were matched by age and sex ratio. Dark-adapted pupil size (<1 lx) was first measured. Then steady-state pupil size was measured during exposure to five lighting conditions (10 lx, 100 lx, 1000 lx, 3000 lx, 6000 lx in the vertical direction at eye level). RESULTS: Significant interaction between the genotype of I394T (TT versus TC+CC) and luminance levels was found in pupil size. Under high illuminance levels (1000 lx, 3000 lx and 6000 lx), pupil sizes in subjects with the C allele were significantly smaller than those in subjects with the TT genotype. On the other hand, pupil size in subjects with the C allele under low illuminance (<1 lx) was significantly larger than that in subjects with the TT genotype. Percentages of pupil constriction under high illuminance levels were significantly greater in subjects with the C allele than in subjects with the TT genotype. CONCLUSIONS: Human melanopsin gene polymorphism I394T interacted with irradiance in association with pupil size. This is the first evidence suggesting a functional connection between melanopsin gene polymorphism and pupillary light response as an index of non-image forming response to light.
  • Event-related potentials reveal subliminal processing of beautiful images
    Sang-il Lee, Satoko Kamiya, Kazuyuki Anekawa, Ryosuke Omura, Takayuki Niahimura, Yeon-kyu Kim, Shigeki Watanuki
    Korean Society for Emotion & Sensibility 163 - 166 2009年05月 [査読無し][通常論文]
  • Physiological Responses Induced by Pleasant and Unpleasant Film Stimuli
    Yeon-kyu Kim, Yoh Hirata, Yuki Motomura, Ryosuke Omura, Sang-il Lee, Takayuki Nishimura, Shigeki Watanuki
    Korean Society for Emotion & Sensibility 141 - 144 2009年 [査読無し][通常論文]
  • キム ヨンキュ, 西村 貴孝, 李 相逸, 綿貫 茂喜
    日本生理人類学会誌 14 2 67 - 74 2009年 [査読有り][通常論文]

講演・口頭発表等

  • Controlling Lighting for Night Shift Workers  [招待講演]
    Sang-il Lee, Shigekazu Higuchi
    The 14th International Congress of Physiological Anthropology 2019 2019年09月 口頭発表(招待・特別)
  • 李相逸, 桶谷康祐
    日本生理人類学会第79回大会 2019年06月 ポスター発表
  • 光の非視覚的作用の研究 -世界の動向-  [通常講演]
    李 相逸
    第1回光と生体リズム研究部会 2018年12月 公開講演,セミナー,チュートリアル,講習,講義等
  • Effects of Short Exposure to Different Light Illuminance in Regular Rest Breaks on Melatonin Suppression and Sleepiness during Simulated Night Work  [通常講演]
    Sang-il Lee, Saki Kinoshita, Anna Noguchi, Taisuke Eto, Kaho Maeda, Yasuhiro Awata, Shigekazu Higuchi
    24th Congress of the European Sleep Research Society 2018年09月 ポスター発表
  • 夜勤仮眠中の漸増光照射とメラトニン分泌の関係  [招待講演]
    李相逸, 樋口重和
    日本睡眠学会第43回定期学術集会 2018年07月 シンポジウム・ワークショップパネル(指名)
  • 夜勤中の光によるメラトニン分泌抑制や概日リズムの位相後退をいかに防ぐか  [招待講演]
    李相逸, 樋口重和
    第24回日本時間生物学会学術大会 2017年10月 シンポジウム・ワークショップパネル(指名)
  • Sang-il Lee, Kota Takeoka, Chie Tarumizu, Sayuri Matsuoka, Shinobu Yasuo, Shigekazu Higuchi
    2017 SSHB - IAPA Joint Symposium 2017年09月 ポスター発表
  • Light-induced melatonin suppression in children under exposure to different color temperature of lights at night  [通常講演]
    Sang-il LEE, Kouhei MATSUMORI, Kana NISHIMURA, Humin XU, Takuya TSUYAMA, Shigekazu HIGUCHI
    23rd Congress of the European Sleep Research Society 2016年09月 ポスター発表
  • Delayed circadian phase is associated with late sleep habits in Japanese children  [通常講演]
    Sang-il Lee, Huimin Xu, Kana Nishimura, Takuya Tsuyama, Shigekazu Higuchi
    20th Congress of the European Anthropological Association 2016年08月 ポスター発表
  • LED照明の色温度と年齢の違いが夜間のメラトニン分泌と眠気に及ぼす影響  [通常講演]
    李相逸, 松森孝平, 西村佳菜, 許慧敏, 津山卓也, 樋口重和
    日本睡眠学会第41回定期学術集会 2016年07月 ポスター発表
  • 色光に対する非視覚的作用と心理的作用の関係  [通常講演]
    李相逸, 落合将太郎, 樋口重和
    日本視覚学会2016年冬季大会 2016年01月 口頭発表(一般)
  • A Proposal to Simplify the Estimation of Characteristic Spectral Responses of Melanopsin to Light Wavelengths  [通常講演]
    Sang-il Lee, Kouhei Matsumori, Shigekazu Higuchi
    The 12th International Congress of Physiological Anthropology 2015年10月 ポスター発表
  • ヒトのメラノプシン遺伝子の一塩基多型I394T と光波長に対する反応特性の関係  [通常講演]
    李相逸, 樋口重和
    第21回日本時間生物学会学術大会 2014年11月 ポスター発表
  • Association between I394T single nucleotide polymorphism of human melanopsin gene and spectral sensitivity of melanopsin  [通常講演]
    Sang-il Lee, Shigekazu Higuchi
    JSC International Symposium 2014年11月 ポスター発表
  • ヒトのメラノプシン遺伝子多型の一塩基多型と睡眠習慣-就寝時刻と覚醒時刻について  [通常講演]
    李相逸, 樋口重和
    日本睡眠学会第39回定期学術集会 2014年07月 ポスター発表
  • The I394T single nucleotide polymorphism of human melanopsin gene is associated with spectral sensitivity of melanopsin  [通常講演]
    Sang-il Lee, Akiko Hida, Shigekazu Higuchi
    26th Society for Light Treatment and Biological Rhythms (SLTBR) 2014年06月 ポスター発表
  • A missense I394T Variant in Human Melanopsin Gene is Associated with Non-Image Forming Responses, including Pupillary Light Reflex and Sleep/Wake Timing  [招待講演]
    Sang-il Lee, Shigekazu Higuchi
    The 1st Asian Conference on Ergonomics and Design (ACED) 2014年05月 口頭発表(一般)
  • Association between genetic polymorphism of melanopsin photoreceptor and sleep/wake timing  [招待講演]
    Sang-il Lee, Shigekazu Higuchi
    The International union of anthropological and ethnological sciences (IUAES) 2014年05月 シンポジウム・ワークショップパネル(指名)
  • メラノプシン遺伝子多型I394Tと睡眠習慣の関係  [通常講演]
    李相逸, 肥田昌子, 北村真吾, 稲見香, 三島和夫, 樋口重和
    第69回日本生理人類学会 2013年10月 ポスター発表
  • Effects of wavelength of light on association between melanopsin gene polymorphism I394T and pupillary light response  [通常講演]
    Sang-il Lee, Shigekazu Higuchi, Akiko Hida, Kazuo Mishima, Sei-ichi Tsujimura, Takeshi Morita
    XIII Congress of the European Biological Rhythms Society 2013年08月 ポスター発表
  • メラノプシン遺伝子多型(I394T)と瞳孔の対光反応の関係:光の強度と色光の影響  [通常講演]
    李相逸, 樋口重和, 西剛史, 肥田昌子, 三島和夫, 辻村誠一, 森田健, 稲見香
    第68回日本生理人類学会 2013年06月 口頭発表(一般)
  • メラノプシン遺伝子多型と瞳孔の対光反応の関係  [通常講演]
    李相逸, 福田知美, 原田和樹, 肥田昌子, 三島和夫, 樋口重和
    第19回日本時間生物学会 2012年09月 ポスター発表
  • Event-related potentials reveal subliminal processing of beautiful images  [通常講演]
    Sang-il Lee, Satoko Kamiya, Kazuyuki Anekawa, Ryosuke Omura, Takayuki Niahimura, Yeon-kyu Kim, Shigeki Watanuki
    Korean Society for Emotion & Sensibility 2009年05月 ポスター発表

受賞

  • 2016年07月 日本睡眠学会 日本睡眠学会第41回定期学術集会ベストプレゼンテーション賞
     
    受賞者: 李 相逸
  • 2014年06月 日本生理人類学会 日本生理人類学会奨励賞
     
    受賞者: 李 相逸
  • 2013年10月 日本生理人類学会 日本生理人類学会第68回大会優秀発表賞
     
    受賞者: 李 相逸
  • 2012年09月 日本時間生物学会 日本時間生物学会第19回大会優秀ポスター賞
     
    受賞者: 李 相逸

共同研究・競争的資金等の研究課題

  • 網膜の光感度の断続的な回復が概日リズムの光同調に及ぼす影響
    日本学術振興会:科学研究費助成事業 基盤研究(C)
    研究期間 : 2020年04月 -2023年03月 
    代表者 : 李 相逸
  • ヒトの概日リズムと覚醒度を考慮した室内環境の検討:照明と室内配色の関係
    日本学術振興会:科学研究費助成事業 若手研究
    研究期間 : 2018年04月 -2020年03月 
    代表者 : 李 相逸
     
    室内の光環境として、照明からの光以外にも、壁などから反射される光(反射光)も含まれる.本研究は、照明と壁の関係によって変わる室内光環境が、ヒトの非視覚的作用(メラトニンの光抑制、概日リズムの位相変動、瞳孔の対光反射など)に及ぼす影響について検討することを目的にしている. 今年度(H30)は予備実験として、壁の配色と距離の違いによる反射光の明るさや波長成分の違いがヒトの瞳孔の対光反射に及ぼす影響について調べた。実験はクロスオーバーデザインで、3色の壁(同素材の白、青、ピンク)と4つの距離(20、40、60、80cm)における瞳孔面積を測定した。その結果、白壁条件時の瞳孔の縮瞳率(%)は、すべての距離において青壁やピンク壁時より有意に高く、その差は距離が遠いほど顕著であった。青壁とピンク壁の間では縮瞳率に有意な差は認められなかった.まだ限られた条件での結果ではあるが、H30年度の研究より、壁の配色と距離によって異なる室内の光環境(波長成分や明るさ)を数値で示し、壁の有無や配色が室内光環境に深く関わっていることを明らかにした.また、瞳孔反射の結果より、夜の光環境を考えるにあたって、白壁の使用は明るさ感を増すには有利であるが、ヒトの生理的機能に余分な刺激を与えることが明らかになった。一方で、青壁はピンク壁に比べて短波長光を多く反射するが、瞳孔反射においてピンク壁以上の影響は与えないことが確認できた.
  • L-セリン摂取が概日リズムの光同調に及ぼす影響
    日本学術振興会:科学研究費助成事業 若手研究(B)
    研究期間 : 2016年04月 -2018年03月 
    代表者 : 李 相逸
     
    本研究では実生活におけるL-セリンの複数回摂取が概日リズム、睡眠習慣に与える影響について明らかにすることを目的とした。健康な男女大学生35名(男:18人女:17人)を対象に4週間のフィールド実験を行った。実験は、L-セリン摂取群とプラセボ摂取群に分けて、二重盲検で実施した。実験1週目(摂取前)から3週目(摂取2週目)のメラトニン分泌開始時刻(概日リズムの位相を示す指標)の差を両群比較したところ有意な違いはなかったが、L-セリン群においてのみ睡眠習慣に対する相対的な前進が確認された。興味深いことに、冬のデータではプラセボ群に比べてL-セリン群のリズムが前進していた。

教育活動情報

主要な担当授業

  • 環境生理学
    開講年度 : 2018年
    課程区分 : 学士課程
    開講学部 : 工学部
    キーワード : 環境適応能 ホメオスタシス 体温調節 呼吸 循環 エネルギー代謝 筋肉 光環境 睡眠 生体リズム

大学運営

委員歴

  • 2017年04月 - 現在   日本生理人類学会   評議員


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