SEARCH

Search Details

Suzuki Keisuke

Center for Human Nature Artificial Intelligence and NeuroscienceAssociate Professor
Research Center of Mathematics for Social CreativityAssociate Professor

Keisuke Suzuki obtained his Ph.D degree on the subject of artificial life from the University of Tokyo in 2007. He stayed as a research fellow in RIKEN Brain Science Institute, where he carried out research into human cognitive functions in virtual reality environments (2008-2011). He developed a novel virtual reality system called SR (Substitutional Reality) with his colleagues. In this system, people believe that they are experiencing real-world scenes, even though they are actually experiencing pre-recorded scenes. After leaving Japan, he worked as a research fellow at the Sackler Centre for Consciousness Science in University of Sussex, UK (2011-2021), where he developed various virtual reality platforms for the experimental study of embodied self-consciousness. In 2021, he joined the Center for Human Nature, Artificial Intelligence, and Neuroscience (CHAIN) in Hokkaido University, Japan as a specially appointed lecturer to continue his work in embodied cognition and conscious presence.

One of Keisuke's key research focuses is about conscious presence; i.e. the subjective feeling of being "here and now". The sense of presence is one of the important aspects of our subjective conscious experience, but its underlying neural mechanisms remain poorly understood. His approach builds on state-of-the-art virtual reality to experimentally manipulate the bodily and mental states of mind, which is complimented by theoretical modeling.

Researcher basic information

■ Degree
  • Ph.D., The University of Tokyo, Mar. 2007
■ URL
researchmap URLホームページURL■ Various IDs
ORCID IDJ-Global ID■ Research Keywords and Fields
Research Keyword
  • Sense of Agency
  • Sense of Body Ownership
  • Altered State of Consciousness
  • Embodied Cognition
  • Visual Hallucination
  • Artificial Life
  • Consciousness
  • Virtual Reality
Research Field
  • Informatics, Human interface and interaction, Virtual Reality
  • Humanities & Social Sciences, Cognitive science, Cognitive Psychology, Embodied Cognitive Science

Career

■ Career
Career
  • Apr. 2024 - Present
    Hokkaido University, Center for Human Nature, Artificial Intelligence and Neuroscience, Associate Professor, Japan
  • Mar. 2021 - Mar. 2024
    Hokkaido University, Center for Human Nature, Artificial Intelligence and Neuroscience, Specially Appointed Lecturer, Japan
  • May 2018 - Feb. 2021
    University of Sussex, Sackler Centre for Consciousness Science, Research Fellow, United Kingdom
  • Sep. 2011 - Apr. 2018
    University of Sussex, Sackler Centre for Consciousness Science, Postdoctral Fellowship, United Kingdom
  • Mar. 2008 - May 2011
    RIKEN, Brain Science Institute, Research Scientist, Japan
Educational Background
  • Apr. 2003 - Mar. 2007, The University of Tokyo, Graduate School of Arts and Sciences, Department of Multi-Disciplinary Sciences,General Systems Sciences, Japan
  • Apr. 2001 - Mar. 2003, The University of Tokyo, Graduate School of Arts and Sciences, Department of Multi-Disciplinary Sciences,General Systems Sciences, Japan
  • Apr. 1997 - Mar. 2001, Sophia University, Department of Physics, Japan
Committee Memberships
  • Nov. 2023 - Oct. 2027
    ISAL - International Society for Artificial Life, Board of Directors (Membership Chair), Society
  • Mar. 2023 - Jul. 2023
    The 2023 Conference on Artificial Life, Sapporo, Japan, programme commitee, Society
  • Aug. 2022 - Jul. 2023
    The 2023 Conference on Artificial Life, Sapporo, Japan, Vice Cheer, Society
  • Mar. 2021 - Jul. 2021
    The 2021 Conference on Artificial Life, in Prague, Czech Republic, Program Committee Member, Society
  • Mar. 2020 - Jul. 2020
    The 2020 Conference on Artificial Life, Online Virtual Conference, Canada, Program Committee Member, Society
  • Mar. 2019 - Jul. 2019
    The 2019 Conference on Artificial Life in Newcastle, UK, Program Committee Member, Society
  • Mar. 2018 - Jul. 2018
    The 2018 Conference on Artificial Life, in Tokyo, Japan, Program Committee Member, Society
  • Mar. 2016 - Jul. 2016
    The 15th International Conference on the Synthesis and Simulation of Living Systems, in Cancún, Mexico, Program Committee Member, Society

Research activity information

■ Awards
  • May 2014, Lorentz Center workshop "Perspectives on Human Probabilistic Inference" at Leiden, the Netherland, The best poster award
    Rubber Hand Illusion Induced by Cardio-Visual Feedback
    Keisuke Suzuki;Sarah N. Garfinkel;Hugo D. Critchley;Anil K. Seth
  • May 2011, INTEREACTION 2011 at Tokyo, Japan, Interactive Presentation Award
    Substitutional Reality: VR system for experiencing "Here and Now"
    Keisuke Suzuki, Sohei Wakisaka, Naotaka Fujii
■ Papers
  • Beyond the reducing valve: towards a computational neurophenomenology of altered states via deep neural networks
    Keisuke Suzuki
    Frontiers in Psychology, 17, Frontiers Media SA, 20 May 2026, [Lead author, Last author, Corresponding author]
    English, Scientific journal, Altered states of consciousness, including hallucinations, psychedelic experiences, and ego dissolution, differ qualitatively, yet no unified computational framework describes what varies and along which dimensions. Computational phenomenology (CP) has emerged as a promising bridge between first-person experience and computational models, yet current formalisations rely predominantly on the free energy principle (FEP). This paper proposes the C × G × D framework, drawing on three functional roles in deep neural networks: a Classifier (C) that extracts features from sensory input, a Generator (G) that synthesises internal representations, and a Discriminator (D) that judges whether a representation originates externally or internally. Phenomenological differences across altered states are redescribed as variations in the objective functions, constraints, and thresholds of these components. The framework reformulates Huxley’s ‘reducing valve’ metaphor: relaxation of C’s constraint exposes normally hidden ‘effective causes,’ producing psychedelic geometric patterns; G’s prior governs hallucinatory veridicality; and D instantiates Perceptual Reality Monitoring. Three hallucination mechanisms—psychedelic, neurodegenerative, and schizophrenia-type—are predicted from distinct parameter configurations. Testable hypotheses derived from iterative-optimisation psychophysics and an extension to ego dissolution are presented. By foregrounding the plurality of objective functions and architectures, the C × G × D framework complements FEP-centred CP and provides a scaffold for translating phenomenology into experimentally manipulable variables., 51596714
  • Intentional binding reflects pair dynamics and sense of agency in embodied joint action in human-human dyads but not in human-computer dyads
    Felix Woolford; Keisuke Suzuki
    Frontiers in Psychology, 17, Frontiers Media SA, 12 Mar. 2026, [Last author, Corresponding author]
    English, Scientific journal, Intentional binding has been proposed as an implicit measure of shared sense of agency in joint action, yet it remains unclear whether it distinguishes between individual and different forms of joint control. We compared intentional binding across individual action, human-computer joint control, and human-human joint control in a physically coupled button-pressing task using haptic feedback devices. Binding magnitude was analyzed both intra-subjectively across conditions and inter-subjectively in relation to reported sense of agency and movement dynamics. Contrary to predictions from the we agency literature, we found no intra-subjective main effect of partner type on binding magnitude. However, within human-human dyads, participants who reported a stronger sense of agency exhibited stronger binding effects, corresponding to emergent leader-follower dynamics in their movement trajectories. No comparable relationship was observed in human-computer interactions. These findings suggest that temporal binding effects primarily reflect sensorimotor predictability rather than intentionality or social context per se. While binding alone does not provide a sufficient marker of “human-like” agency in artificial systems, it may reflect the distribution of predictive control within a dyad and thus serve as a quantifiable signature of how effectively partners co-regulate their actions., 33902486
  • Multisensory integration in chaotic networks
    Adam Ponzi; Keisuke Suzuki
    Neural Networks, 191, 107766, 107766, Elsevier BV, Nov. 2025, [Peer-reviewed], [Last author]
    English, Scientific journal, 33902486
  • Investigating the role of sensorimotor spatial dependencies in shaping conscious access to virtual 3D objects
    Paweł Motyka; David J. Schwartzman; Anil K. Seth; Keisuke Suzuki
    Consciousness and Cognition, 135, 103934, 103934, Elsevier BV, Oct. 2025, [Peer-reviewed], [Last author]
    English, Scientific journal, 33902486
  • Real-time fMRI neurofeedback boosts heartbeat perception by modulating insula activation pattern during interoceptive attention
    Yusuke Haruki; Yuxiang Yang; Keisuke Suzuki; Hiroshi Imamizu; Kenji Ogawa
    Imaging Neuroscience, MIT Press, 21 Aug. 2025, [Peer-reviewed]
    English, Scientific journal, Abstract

    Real-time fMRI neurofeedback (NF) has emerged as a promising method for enabling individuals to modulate specific brain regions and, consequently, their behavioural outcomes. This study examined whether the NF targeting the right insula could improve heartbeat perception ability and influence emotional response to negatively valenced stimuli, by training participants to modulate the brain activation associated with interoceptive (heartbeat-focused) and exteroceptive (visual-focused) attention. Fifty-four participants underwent a single ~40-minute NF session with contingent (NF group, n = 28) or non-contingent (Sham group, n = 26) feedback, with heartbeat perception and emotional appraisal assessed pre- and post-training. The NF group demonstrated significant improvements in heartbeat perception, with individual learning effects in neuromodulation predicting the behavioural gains. However, group-level NF scores did not differ significantly, likely reflecting variability in learnability. Despite improvements in heartbeat perception, NF training did not modulate emotional responses at either the behavioural or neural level, suggesting that targeting the insula alone is insufficient to alter affective processing within a single session. These findings provide evidence that NF can enhance heartbeat perception through targeted neuromodulation in the insular cortex., 33902486
  • Active inference for action-unaware agents
    Filippo Torresan; Keisuke Suzuki; Ryota Kanai; Manuel Baltieri
    arXiv, Aug. 2025
    English, Scientific journal, 33902486
  • The Japanese version of the Phenomenological Control Scale
    Shu Imaizumi; Keisuke Suzuki
    Neuroseicne of Consciousness, 2025, 1, Center for Open Science, 22 May 2025, [Peer-reviewed], [Last author, Corresponding author]
    English, Scientific journal,

    People possess the capacity for phenomenological control that enables them to align their perceptual experiences with their intentions and goals. The Phenomenological Control Scale was developed to measure this trait, and we developed and validated a Japanese version of this scale (PCS-J) based on preregistered online surveys (n = 261; retest n = 152). The PCS-J demonstrated sufficient internal consistency and test-retest reliability. Given the known association between hypnotic susceptibility and positive schizotypy, the convergent validity of the PCS-J was supported by a weak positive correlation with positive schizotypy. The discriminant validity of the PCS-J was demonstrated by the absence of a correlation with negative schizotypy. The PCS-J would be useful for research on perception, phenomenological control, and their individual differences in Japanese samples as well as for intercultural studies.

    , 33902486
  • The Vagus Nerve as a Gateway to Body Ownership: taVNS Reduces Susceptibility to a Virtual Version of the Cardiac and Tactile Rubber Hand Illusion
    Alisha Vabba; Keisuke Suzuki; Milica Doric; Tim J. Möller; Sarah Garfinkel; Hugo Critchley
    Psychophysiology, 62, 3, Wiley, 17 Mar. 2025, [Peer-reviewed]
    English, Scientific journal, ABSTRACT

    Transcutaneous auricular vagus nerve stimulation (taVNS) has been shown to influence cognitive and emotional function and enhance interoceptive awareness. This study investigates if taVNS effects extend to the experience of body ownership, as measured via susceptibility to the rubber hand illusion (RHI) in a virtual reality setting. The experiment involved 27 participants who underwent real and sham stimulation in two separate sessions while experiencing synchronous or asynchronous visuo‐cardiac and visuo‐tactile feedback on a virtual arm in place of their own. Results indicated that active compared to sham taVNS decreased sensitivity to the illusion in both cardiac and tactile trials. Specifically, a greater proprioceptive drift difference (PDD) toward the rubber hand was observed for synchronous compared to asynchronous trials only during sham (t(26) = −4.58, pbonf < 0.001) but not during active (pbonf = 1.00) stimulation. A similar pattern was also observed for subjective ownership, where synchronous trials led to greater subjective ownership than asynchronous trials only during sham (t(26) = −3.52, pbonf = 0.010) but not during active (pbonf = 1.00) stimulation. These findings suggest that stimulation might enhance body ownership, making individuals more attuned to their real bodily signals and less susceptible to bodily illusions. Additionally, physiological measures such as heart rate (HR), heart rate variability (HRV), and skin sympathetic nervous activity (SKNA) were assessed to explore the autonomic effects of taVNS. We observed a decrease in HR during active stimulation (t(26) = 4.30, pbonf < 0.001), and an increase in SKNA during both sham (t(26) = −4.40, pbonf < 0.001) and active stimulation (t(26) = −4.85, pbonf < 0.002). These findings contribute to the understanding of the vagus nerve's role in integrating visceral and somatosensory signals, with implications for clinical applications in conditions characterized by altered interoception and body ownership., 33902486
  • Attentional bias towards smartphone stimuli is associated with decreased interoceptive awareness and increased physiological reactivity
    Yusuke Haruki; Katsunori Miyahara; Kenji Ogawa; Keisuke Suzuki
    Communications Psychology, 17 Mar. 2025
    Scientific journal
  • Modelling Phenomenological Differences in Aetiologically Distinct Visual Hallucinations Using Deep Neural Networks
    Keisuke Suzuki; Anil K. Seth; David J. Schwartzman
    Frontier in Human Neuroscience, 17, Cold Spring Harbor Laboratory, 03 Jan. 2024, [Lead author]
    English, Scientific journal, Abstract

    Visual hallucinations (VHs) are perceptions of objects or events in the absence of the sensory stimulation that would normally support such perceptions. Although all VHs share this core characteristic, there are substantial phenomenological differences between VHs that have different aetiologies, such as those arising from neurological conditions, visual loss, or psychedelic compounds. Here, we examine the potential mechanistic basis of these differences by leveraging recent advances in visualising the learned representations of a coupled classifier and generative deep neural network – an approach we call ‘computational (neuro)phenomenology’. Examining three aetiologically distinct populations in which VHs occur - neurological conditions (Parkinson’s Disease and Lewy Body Dementia), visual loss (Charles Bonnet Syndrome, CBS), and psychedelics - we identify three dimensions relevant to distinguishing these classes of VHs: realism (veridicality), dependence on sensory input (spontaneity), and complexity. By selectively tuning the parameters of the visualisation algorithm to reflect influence along each of these phenomenological dimensions we were able to generate ‘synthetic VHs’ that were characteristic of the VHs experienced by each aetiology. We verified the validity of this approach experimentally in two studies that examined the phenomenology of VHs in neurological and CBS patients, and in people with recent psychedelic experience. These studies confirmed the existence of phenomenological differences across these three dimensions between groups, and crucially, found that the appropriate synthetic VHs were representative of each group’s hallucinatory phenomenology. Together, our findings highlight the phenomenological diversity of VHs associated with distinct causal factors and demonstrate how a neural network model of visual phenomenology can successfully capture the distinctive visual characteristics of hallucinatory experience.
  • 自由エネルギー原理,能動的視覚,サリエンス
    吉田 正俊; 宮園 健吾; 西尾 慶之; 山下 祐一; 鈴木 啓介
    人工知能, 38, 6, 787, 795, 一般社団法人 人工知能学会, 01 Nov. 2023
    Japanese
  • Hybrid Life: Integrating biological, artificial, and cognitive systems
    Manuel Baltieri; Hiroyuki Iizuka; Olaf Witkowski; Lana Sinapayen; Keisuke Suzuki
    WIREs Cognitive Science, Wiley, 05 Jul. 2023
    Scientific journal, Abstract

    Artificial life is a research field studying what processes and properties define life, based on a multidisciplinary approach spanning the physical, natural, and computational sciences. Artificial life aims to foster a comprehensive study of life beyond “life as we know it” and toward “life as it could be,” with theoretical, synthetic, and empirical models of the fundamental properties of living systems. While still a relatively young field, artificial life has flourished as an environment for researchers with different backgrounds, welcoming ideas, and contributions from a wide range of subjects. Hybrid Life brings our attention to some of the most recent developments within the artificial life community, rooted in more traditional artificial life studies but looking at new challenges emerging from interactions with other fields. Hybrid Life aims to cover studies that can lead to an understanding, from first principles, of what systems are and how biological and artificial systems can interact and integrate to form new kinds of hybrid (living) systems, individuals, and societies. To do so, it focuses on three complementary perspectives: theories of systems and agents, hybrid augmentation, and hybrid interaction. Theories of systems and agents are used to define systems, how they differ (e.g., biological or artificial, autonomous, or nonautonomous), and how multiple systems relate in order to form new hybrid systems. Hybrid augmentation focuses on implementations of systems so tightly connected that they act as a single, integrated one. Hybrid interaction is centered around interactions within a heterogeneous group of distinct living and nonliving systems. After discussing some of the major sources of inspiration for these themes, we will focus on an overview of the works that appeared in Hybrid Life special sessions, hosted by the annual Artificial Life Conference between 2018 and 2022.

    This article is categorized under:Neuroscience > Cognition

    Philosophy > Artificial Intelligence

    Computer Science and Robotics > Robotics
  • Sensorimotor coupling but not visual perspective modulates perceived time during voluntary action
    Shu Imaizumi; Giuseppe Lai; Anil K. Seth; Keisuke Suzuki
    PsyArXiv, Center for Open Science, May 2023

    Voluntary actions are associated with both a sense of agency of the action, and with distortions of perceived duration. The present study conducted four pre-registered experiments using virtual reality to investigate whether and how perceived duration and sense of agency during voluntary action are modulated by sensorimotor coupling and visual perspective. Participants performed hand movements while observing a virtual hand moving (a)synchronously with their own hand, estimated the duration of the movements, and rated their sense of agency over the movements. Pre-registered analyses revealed that sensorimotor coupling provided by synchronous visual feedback dilates perceived duration, compared to delayed feedback or pre-recorded feedback of other’s movements (sensorimotor decoupling) in Experiments 1 and 2. Visual perspective (Experiments 1 and 2), anatomical configuration (Experiment 3), and visual eccentricity of the virtual hand (Experiment 4) did not influence duration perception. Sense of agency was also modulated by sensorimotor coupling, suggesting that sensorimotor coupling could be a shared foundation for duration perception and sense of agency. Our findings and the virtual-reality technique we developed shed new light on how humans experience time and self-agency during voluntary action.

  • Awareness of cardiac/gastric interoception and its individual differences modulate the neural dynamics
    Haruki Yusuke; Suzuki Keisuke; Ogawa Kenji
    Cognitive Studies: Bulletin of the Japanese Cognitive Science Society, 30, 1, 108, 111, Japanese Cognitive Science Society, 01 Mar. 2023
    English
  • A neurophenomenological approach to non-ordinary states of consciousness: hypnosis, meditation, and psychedelics
    Christopher Timmermann; Prisca R. Bauer; Olivia Gosseries; Audrey Vanhaudenhuyse; Franz Vollenweider; Steven Laureys; Tania Singer; Elena Antonova; Antoine Lutz; Mind and Life Europe (MLE) ENCECON Research Group (incl. Suzuki K.)
    Trends in Cognitive Sciences, 27, 2, 139, 159, Elsevier BV, Feb. 2023
    English, Scientific journal
  • Using extended reality to study the experience of presence
    K Suzuki; A Mariola; DJ Schwartzman; AK Seth
    Current Topics in Behavioral Neurosciences, 1, 31, Jan. 2023
    English, In book
  • Who tailors the blanket?
    Keisuke Suzuki; Katsunori Miyahara; Kengo Miyazono
    Behavioral and Brain Sciences, 45, e213, Center for Open Science, 27 Sep. 2022, [Peer-reviewed], [Lead author]
    English, Scientific journal,

    The gap between the Markov blanket and ontological boundaries arises from the former’s inability to capture the dynamic process through which biological and cognitive agents actively generate their own boundaries with the environment. Active inference in the FEP framework presupposes the existence of a Markov blanket, but it is not a process that actively generates the latter.

  • Cognitive Science of Embodied Self-Consciousness
    Keisuke Suzuki; Katsunori Miyahara
    生体の科学, 73, 1, Feb. 2022, [Invited], [Lead author, Corresponding author]
    Japanese, Scientific journal
  • Gradient Climbing Neural Cellular Automata
    Shuto Kuriyama; Wataru Noguchi; Hiroyuki Iizuka; Keisuke Suzuki; Masahito Yamamoto
    The 2022 Conference on Artificial Life, MIT Press, 2022
    International conference proceedings
  • Virtual Reality and "Mind-as-it-could-be"
    Keisuke Suzuki
    人工知能, 37, 1, 27, 34, Jan. 2022, [Peer-reviewed], [Invited], [Lead author, Last author, Corresponding author]
    Japanese, Scientific journal
  • More than meets the heart: systolic amplification of different emotional faces is task dependent.
    Mateo Leganes-Fonteneau; Jennifer F Buckman; Keisuke Suzuki; Anthony Pawlak; Marsha E Bates
    Cognition & emotion, 35, 2, 400, 408, Mar. 2021, [Peer-reviewed], [International Magazine]
    English, Scientific journal, Interoceptive processes emanating from baroreceptor signals support emotional functioning. Previous research suggests a unique link to fear: fearful faces, presented in synchrony with systolic baroreceptor firing draw more attention and are rated as more intense than those presented at diastole. This study examines whether this effect is unique to fearful faces or can be observed in other emotional faces. Participants (n = 71) completed an emotional visual search task (VST) in which fearful, happy, disgust and sad faces were presented during systolic and diastolic phases of the cardiac cycle. Visual search accuracy and emotion detection accuracy and latency were recorded, followed by a subjective intensity task. A series of interactions between emotion and cardiac phase were observed. Visual search accuracy for happy and disgust faces was greater at systole than diastole; the opposite was found for fearful faces. Fearful and happy faces were perceived as more intense at systole. Previous research proposed that cardiac signalling has specific effects on the attention and intensity ratings for fearful faces. Results from the present tasks suggest these effects are more generalised and raise the possibility that interoceptive signals amplify emotional superiority effects dependent on the task employed.
  • Sensorimotor contingency modulates breakthrough of virtual 3D objects during a breaking continuous flash suppression paradigm
    Keisuke Suzuki; David J Schwartzman; Rafael Augusto; Anil K Seth
    Cognition, 187, 95, 107, Elsevier BV, Jun. 2019, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Intentional Binding Without Intentional Action
    Keisuke Suzuki
    Psychological Science, 30, 6, 842, 853, SAGE Publications, Jun. 2019, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal, The experience of authorship over one’s actions and their consequences—sense of agency—is a fundamental aspect of conscious experience. In recent years, it has become common to use intentional binding as an implicit measure of the sense of agency. However, it remains contentious whether reported intentional-binding effects indicate the role of intention-related information in perception or merely represent a strong case of multisensory causal binding. Here, we used a novel virtual-reality setup to demonstrate identical magnitude-binding effects in both the presence and complete absence of intentional action, when perceptual stimuli were matched for temporal and spatial information. Our results demonstrate that intentional-binding-like effects are most simply accounted for by multisensory causal binding without necessarily being related to intention or agency. Future studies that relate binding effects to agency must provide evidence for effects beyond that expected for multisensory causal binding by itself.
  • Sensorimotor contingency modulates visual awareness of virtual 3D objects
    Keisuke Suzuki; David J. Schwartzman; Rafael Augusto; Anil K. Seth
    The 2019 Conference on Artificial Life, 2019, [Peer-reviewed], [Lead author, Corresponding author]
    English, International conference proceedings
  • Hallucination Machine: Simulating Altered Perceptual Phenomenology with a Deep-Dream Virtual Reality platform
    Keisuke Suzuki; Warrick Roseboom; David J. Schwartzman; Anil K. Seth
    The 2018 Conference on Artificial Life, 2018, [Peer-reviewed], [Lead author, Corresponding author]
    English, International conference proceedings
  • A Deep-Dream Virtual Reality Platform for Studying Altered Perceptual Phenomenology
    Keisuke Suzuki
    Scientific Reports, 7, 1, Springer Science and Business Media LLC, Dec. 2017, [Peer-reviewed], [Lead author, Corresponding author]
    English, Scientific journal
  • Predictive coding accounts of shared representations in parieto-insular networks
    Hiroaki Ishida; Keisuke Suzuki; Laura Clara Grandi
    Neuropsychologia, 70, 442, 454, Elsevier BV, Apr. 2015, [Peer-reviewed], [Invited], [Lead author]
    English, Scientific journal
  • Knowing your own heart: Distinguishing interoceptive accuracy from interoceptive awareness
    Sarah N Garfinkel; Anil K Seth; Adam B Barre; Keisuke Suzuki; Hugo D Critchley
    Biological Psychology, 104, 65, 74, Elsevier BV, Jan. 2015, [Peer-reviewed]
    English, Scientific journal
  • Multisensory integration across exteroceptive and interoceptive domains modulates self-experience in the rubber-hand illusion
    Keisuke Suzuki
    Neuropsychologia, 51, 13, 2909, 2917, Elsevier BV, Nov. 2013, [Peer-reviewed], [Lead author]
    English, Scientific journal
  • Substitutional Reality System: A Novel Experimental Platform for Experiencing Alternative Reality
    Keisuke Suzuki; Sohei Wakisaka; Naotaka Fujii
    Scientific Reports, 2, 1, Springer Science and Business Media LLC, Dec. 2012, [Peer-reviewed], [Lead author]
    English, Scientific journal
  • Using Human–Computer Interfaces to Investigate ‘Mind-As-It-Could-Be’ from the First-Person Perspective
    Tom Froese; Keisuke Suzuki; Yuta Ogai; Takashi Ikegami
    Cognitive Computation, 4, 3, 365, 382, Sep. 2012, [Peer-reviewed]
    English, Scientific journal
  • AN EXPERIMENTAL APPROACH TO SPEAKERS' PERSPECTIVES IN A PAIRED WALL GAME
    RYOKO UNO; KEISUKE SUZUKI; TAKASHI IKEGAMI
    Advances in Complex Systems, 15, 03n04, 1150020, 1150020, May 2012, [Peer-reviewed]
    English, Scientific journal
  • Adaptability and Homeostasis in the Game of Life interacting with the evolved Cellular Automata
    Keisuke Suzuki; Takashi Ikegami
    Nature-Inspired Computing Design, Development, and Applications, 232, 254, IGI Global, 2012, [Peer-reviewed], [Lead author]
    English, In book, In this paper, the authors study the emergence of homeostasis in a two-layer system of the Game of Life, in which the Game of Life in the first layer couples with another system of cellular automata in the second layer. Homeostasis is defined as a space-time dynamic that regulates the number of cells in state-1 in the Game of Life layer. A genetic algorithm is used to evolve the rules of the second layer to control the pattern of the Game of Life. The authors found that two antagonistic attractors control the numbers of cells in state-1 in the first layer. The homeostasis sustained by these attractors is compared with the homeostatic dynamics observed in Daisy World.
  • An Interoceptive Predictive Coding Model of Conscious Presence
    Keisuke Suzuki
    Frontiers in Psychology, 2, Frontiers Media SA, 2012, [Peer-reviewed]
    English, Scientific journal
  • From artificial life to artificial embodiment: Using human-computer interfaces to investigate the embodied mind 'as-it-could-be' from the first-person perspective
    Tom Froese; Keisuke Suzuki; Sohei Wakisaka; Yuta Ogai; Takashi Ikegami
    AISB 2011: Computing and Philosophy, 43, 50, 2011
    International conference proceedings
  • Substitutional Reality: a VR System to Experience "Here and Now"
    Keisuke Suzuki; Sohei Wakisaka; Naotaka Fujii
    Interaction2012 予稿集, 2011, [Peer-reviewed], [Lead author]
    Japanese
  • An interactive wall game as an evolution of proto language.
    Ryoko Uno; Keisuke Suzuki; Takashi Ikegami
    Advances in Artificial Life: 20th Anniversary Edition - Back to the Origins of Alife(ECAL), 813, 819, MIT Press, 2011, [Peer-reviewed]
    International conference proceedings
  • Language as Autopoiesis - Experimental Approach to Agency in Linguistic Communication.
    Keisuke Suzuki; Ryoko Uno; Takashi Ikegami
    Artificial Life XII: Proceedings of the Twelfth International Conference on the Synthesis and Simulation of Living Systems(ALIFE), 860, 861, MIT Press, 2010, [Peer-reviewed], [Lead author]
    English, International conference proceedings
  • Shapes and Self-Movement in Protocell Systems
    Keisuke Suzuki; Takashi Ikegami
    Artificial Life, 15, 1, 59, 70, MIT Press - Journals, Jan. 2009, [Peer-reviewed], [Lead author], [International Magazine]
    English, Scientific journal, The effect of shapes on self-movement has been studied with an extended model of autopoiesis. Autopoiesis is known as a theory of self-boundary maintenance. In this study, not only the autopoietic generation of the self-boundary, but also the emergence of self-motility, has been examined. As a result of computer simulations, it has been found that different membrane shapes cause different types of self-movement. A kind of chemotaxis has been observed for some shapes. The mechanism of chemotaxis is discussed by studying the internal chemical processes within the shape boundaries.
  • From a homeostatic to a homeodynamic self
    Takashi Ikegami; Keisuke Suzuki
    Biosystems, 91, 2, 388, 400, Elsevier BV, Feb. 2008, [Peer-reviewed], [Lead author], [International Magazine]
    English, Scientific journal, Life as an autonomous homeostatic system is discussed. A mechanism that drives a homeostatic state to an autonomous self-moving state is examined with two computational cell models. The mechanism is met with Ashby's ultrastability, where random parameter searching is activated when a system breaks a viability constraint. Such a random search process is replaced by the membrane shape in the first model and by chaotic population dynamics in the second model. Emergence of sensors, motors and the recursive coupling between them is shown to be a natural outcome of an autonomous homeostatic system.
  • Homeodynamics in the Game of Life
    Keisuke Suzuki; Takashi Ikegami
    Artificial Life XI: Proceedingshe of the Eleventh International Conference on the Synthesis and Simulation of Living Systems(ALIFE), 600, 607, MIT Press, 2008, [Peer-reviewed], [Lead author]
    English, International conference proceedings
  • Morphological dynamics in a proto-cell model
    Keisuke Suzuki; Takashi Ikegami
    PROCEEDINGS OF SICE ANNUAL CONFERENCE, VOLS 1-8, 901, +, 2007
    English, International conference proceedings
  • Spatial-Pattern-Induced Evolution of a Self-Replicating Loop Network
    Keisuke Suzuki; Takashi Ikegami
    Artificial Life, 12, 4, 461, 485, MIT Press - Journals, Oct. 2006, [Peer-reviewed], [Lead author]
    English, Scientific journal, We study a system of self-replicating loops in which interaction rules between individuals allow competition that leads to the formation of a hypercycle-like network. The main feature of the model is the multiple layers of interaction between loops, which lead to both global spatial patterns and local replication. The network of loops manifests itself as a spiral structure from which new kinds of self-replicating loops emerge at the boundaries between different species. In these regions, larger and more complex self-replicating loops live for longer periods of time, managing to self-replicate in spite of their slower replication. Of particular interest is how micro-scale interactions between replicators lead to macro-scale spatial pattern formation, and how these macro-scale patterns in turn perturb the micro-scale replication dynamics.
  • Interaction Based Evolution of Self-Replicating Loop Structures.
    Keisuke Suzuki; Takashi Ikegami
    ECAL 2003: Advances in Artificial Life, 89, 96, Springer, 2003, [Peer-reviewed], [Lead author]
    English, International conference proceedings
■ Other Activities and Achievements
  • 能動的推論:生存と適応の認知理論
    鈴木 啓介, 日本バーチャルリアリティ学会誌, 29, 2, 54, 54, 30 Jun. 2024
    特定非営利活動法人 日本バーチャルリアリティ学会, Japanese
  • Salience as affordance: implications for aberrant salience hypothesis of psychosis from active inference perspective
    Masatoshi Yoshida; Kengo Miyazono; Yoshiyuki Nishio; Yuichi Yamashita; Keisuke Suzuki, Jxiv, 01 Sep. 2022, [Domestic magazines]
    サリエンスという言葉には多義性がある。かたや「知覚サリエンス」では知覚的に周りと比べて目立つという特性を表しており、かたや「動機サリエンス」では物体に情動価を付与させる働きとして捉えられている。本論文では「知覚サリエンス」と「動機サリエンス」との関係を明確にすることを目的とする。このことは精神医学において提唱されているASHを精緻化できるという貢献があることが期待される。精神病症状の異常サリエンス仮説aberrant salience hypothesis of psychosis (ASH)は精神病症状の発達から回復までを整合的に説明したものとして重要性が高い。しかし重要な問題点としてサリエンスという言葉が曖昧だという点がある。そこでこの論文ではまず現状でのサリエンスの定義を明確にするため、知覚サリエンスと動機サリエンスに分けて考える。そしてこのサリエンスをアフォーダンスとして捉える考え方がふたつの方向性 (心理学の哲学と情報理論的な脳のモデル(能動的推論))から支持されることを示す。このような視点からサリエンスを捉え直すことで、サリエンスはepistemic affordance, exploitive affordance, aversive affordanceに分類できる。このようにサリエンスを捉え直すことによって、ASHはより精緻なものとできる。つまりASHにおいては、epistemic affordanceとaversive affordance両方の亢進を考慮する必要がある。ここで前者は対象への接近を動機づけるが、後者は対象の回避を動機づける。このようなコンフリクトが患者を苦しめ、なんらかの認知的な解決を必要とする。このことはASHでの「サリエンスの誤帰属」という言葉からは見えてこない。最後に以上のことと脳部位との対応について議論し、さらに本論文で提案される考えの応用範囲について議論する。, 126–141, Japanese, Technical report
  • バーチャルリアリティと変性意識体験 (特集 バーチャルリアリティの広がり)
    鈴木 啓介, 心理学ワールド, 88, 17, 20, Jan. 2020
    日本心理学会, Japanese
  • Emergence of leader/follower distinction in proto-linguistic communication
    宇野良子; 飯塚博幸; 鈴木啓介, 電子情報通信学会技術研究報告, 112, 176(HCS2012 33-46), 2012
■ Books and other publications
■ Lectures, oral presentations, etc.
■ Syllabus
  • 大学院共通授業科目(教育プログラム):人間知・脳・AI教育プログラム, 2024年, 修士課程, 大学院共通科目
  • 大学院共通授業科目(教育プログラム):人間知・脳・AI教育プログラム, 2024年, 修士課程, 大学院共通科目
  • 大学院共通授業科目(教育プログラム):人間知・脳・AI教育プログラム, 2024年, 修士課程, 大学院共通科目
  • 大学院共通授業科目(教育プログラム):人間知・脳・AI教育プログラム, 2024年, 修士課程, 大学院共通科目
  • 大学院共通授業科目(教育プログラム):人間知・脳・AI教育プログラム, 2024年, 修士課程, 大学院共通科目
  • 大学院共通授業科目(一般科目):複合領域, 2024年, 修士課程, 大学院共通科目
  • 思索と言語, 2024年, 学士課程, 全学教育
■ Research Themes
  • Mesoscopic-level digital brain models for the development of novel therapeutic interventions for mental disorders.
    2024 Integrated Programme for Neuroscience (Individual Priority Research Programmes) 4-1 'Area 4: Development of brain models reproduced in digital space and construction of research infrastructure (digital brain)' Team type B
    Oct. 2024 - Mar. 2029
    Yuichi Yamashita; Maki Iwasaki, Kaike Suzuki
    Japan Agency for Medical Research and Development, Center for Human Nature, Artificial Intelligence and Neuroscience (CHAIN), Hokkaido University, Coinvestigator, 24018895
  • 人と人工エージェントの身体的社会相互作用における感覚運動ダイナミクスの研究
    科学研究費助成事業
    15 Nov. 2024 - 31 Mar. 2027
    鈴木 啓介
    日本学術振興会, 特別研究員奨励費, 北海道大学, 24KF0170
  • Philosophical and Interdisciplinary Studies of Atypical Experience, Belief, and Cognition
    Grants-in-Aid for Scientific Research
    Apr. 2024 - Mar. 2027
    宮園 健吾; ディーツ リチャード; PEEBLES GRAHAM; 吉田 正俊; 山下 祐一; オデイ ジョン; FRISCHHUT Akiko; 鈴木 啓介; 稲荷森 輝一; 西尾 慶之
    Japan Society for the Promotion of Science, Grant-in-Aid for Scientific Research (A), Hokkaido University, 24H00076
  • Cognitive Feelings that Mediate Between Perception and Emotion
    Oct. 2021 - Mar. 2027
    JST CREST, Coinvestigator
  • Sharing and controlling diversity of inner speech using VR technology
    Grants-in-Aid for Scientific Research
    01 Apr. 2024 - 31 Mar. 2026
    鈴木 啓介
    Japan Society for the Promotion of Science, Grant-in-Aid for Transformative Research Areas (A), Hokkaido University, 24H01534
  • Synthetic approach to perceptual illusions using cognitive deep learning model with predictive learning
    Grants-in-Aid for Scientific Research
    01 Apr. 2022 - 31 Mar. 2026
    飯塚 博幸; 山本 雅人; 鈴木 啓介; 野口 渉
    令和5年度は、初年度に開発したボトムアップ処理を中心とする深層学習モデルを拡張し、重ね合わせ原理を用いた感覚統合を行うニューラルネットワークモデルを用いて、感覚入力のトップダウン的解釈を与える認知モデルを開発した。このモデルは、予測符号化理論に基づいており、感覚入力に対する予測と予測誤差の最小化を通じて、感覚情報の統合と解釈を行う。また、ラバーハンド錯覚のシミュレーションモデルを利用し、自分の身体領域を道具の先までに拡張するシミュレーションモデルを構築した。トップダウン的解釈を可能とする認知モデルにより、道具使用時の触感覚を道具不使用時の自分の身体の触感覚へと同等に重ね合わせることに成功した。
    近接領域での手を用いた作業や振る舞いの経験が錯覚に与える影響を調べるため、実環境での、視覚、触覚、運動データの計測準備を進めた。協力者の頭部にカメラを固定し、作業時の画像を視覚データとして取得し、手にはデータグローブを装着し、触覚データと運動データを収集するこれらのデータは、初年度と2年目に行ったシミュレーションと同等の実環境データとして活用される。トップダウン的解釈を可能とする認知モデルとシミュレーション・物理環境において収集したデータを用いて、深層学習で実装した認知モデルを学習させた。ラバーハンド錯覚のシミュレーションにおいて、錯覚を再現することに部分的に成功しており、今後の更なる改善が期待される。
    本研究の成果は、国際学会で発表し、高い評価を得ている。トップダウン的解釈を可能とする認知モデルの開発と、シミュレーションデータを用いた学習手法について、論文を執筆しており、国際学術誌に投稿する準備を行った。令和5年度の研究により、トップダウン的解釈を可能とするモデルの開発とそれらを用いた認知モデルの学習が進展した。
    Japan Society for the Promotion of Science, Grant-in-Aid for Scientific Research (B), Hokkaido University, 23K25163
  • 脳状態可視化に基づく内受容注意モニタリングシステム開発とマインドフルネスへの応用
    科学研究費助成事業 挑戦的研究(萌芽)
    Jun. 2022 - Mar. 2025
    小川 健二
    本研究は、被験者自身の内受容注意に関わる脳活動を可視化して本人にフィードバックすることで、効果的な脳のセルフコントロール・システムを提案する。身体内部からの情報を伝える感覚(内受容感覚)は、特に感情・情動やメンタルヘルス等との関連が示されている。マインドフルネスでは、現在の自分の身体内部感覚(呼吸や心拍)に注意(内受容注意)を向ける (body scan)ことで、外的なストレスへの 効果的な対処を図っており、心理臨床現場でもその効果が実証されている。しかし運動訓練等とは異なり、内受容注意の程度、すなわち自身の脳活動変化を観察することはできないため、どのような方略を取れば効果的であるのかが明確でないという問題点がある。そこで本研究はリアルタイムfMRIを使い、内受容注意時の自身の脳活動を可視化し本人にフィードバックすることで、自身の脳状態を把握した効果的な脳のセルフコントロール・システムの開発と応用を目指す。本提案では脳の島皮質に着目する。島皮質は大脳皮質が島のように内側に折り畳まれている箇所であり、視床から内臓感覚などの身体内部に関する感覚(内受容感覚)を前頭葉へと中継する場所である。またfMRIを使った研究からも、内受容注意により、島皮質の活動が増加する点が明らかになっている。実験では、被験者にMRI装置に横たわった状態で内受容注意課題を行ってもらう。具体的には、軽く目を空けた状態で自分の現在の身体感覚(呼吸や心拍など)に注意を向ける。その際の島皮質の活動をfMRIで計測し、その結果を本人に対してリアルタイムで棒グラフや数値等で分かりやすく視覚呈示でフィードバックする。
    日本学術振興会, 挑戦的研究(萌芽), 北海道大学, Coinvestigator, 22K18643
  • Elucidating the effects of the relationship with digital space on bodily awareness and body diagrams.
    Life Design for Mind and Body Young Research Fund 2023
    Apr. 2023 - Mar. 2024
    Keisuke Suzuki
    Open Innovation Platforms for Industry-academia Co-creation (COI-NEXT), Center for Human Nature, Artificial Intelligence and Neuroscience (CHAIN), Hokkaido University, Principal investigator, Competitive research funding
■ Industrial Property Rights
  • Control device for alternate reality system, alternate reality system, control method for alternate reality system, program, and recording medium
    Patent right, Naotaka Fujii; Sohei Wakisaka; Keisuke Suzuki, RIKEN Institute of Physical and Chemical Research
    特願14/422,109, 15 Jul. 2013
    特開US20150227196A1, 13 Jul. 2015
■ Academic and Social Contribution Activities/Other
Industrial Property Rights
  • ALife 2025
    06 Oct. 2025 - 10 Oct. 2025
    Planning etc, Panel chair etc
    Academic society etc
    Olaf Witkowski, Alyssa Adams
  • Models of Consciousness 2025
    30 Sep. 2025 - 04 Oct. 2025
    Planning etc, Panel chair etc, Review, Peer review
    Academic society etc
    Robert Prentner, Keisuke Suzuki, Joanna Szczotka, Johannes Kleiner, Jonathan Mason, Wanja Wiese, Camilo Miguel Signorelli, Robin Lorenz
  • The 2023 Conference on Artificial Life
    24 Jul. 2023 - 28 Jul. 2023
    Planning etc, Panel chair etc
    Academic society etc
    Hiroyuki Iizuka, Keisuke Suzuki, Ryoko Uno, Luisa Damiano, Nadine Spychala, Miguel Aguilera, Eduardo Izuierdo, Reiji Suzuki, Manuel Baltieri
  • A special session in ALIFE2021: Hybrid Life IV: Approaches to integrate biological, artificial and cognitive systems
    19 Jul. 2021 - 23 Jul. 2021
    Planning etc, Panel chair etc, Peer review
    Academic society etc
    Manuel Baltieri, Keisuke Suzuki, Hiroyuki Iizuka, Olaf Witkowski, Lana Sinapayen
  • A special session in ALIFE2020: Hybrid Life III: Approaches to integrate biological, artificial and cognitive systems
    13 Jul. 2020 - 18 Jul. 2020
    Planning etc, Panel chair etc, Peer review
    Academic society etc
    Manuel Baltieri, Keisuke Suzuki, Hiroyuki Iizuka, Olaf Witkowski, Lana Sinapayen
  • A special session in ALIFE2019: Hybrid Life II: Approaches to integrate biological, artificial and cognitive systems
    29 Jul. 2019 - 02 Aug. 2019
    Planning etc, Panel chair etc, Peer review
    Academic society etc
    Manuel Baltieri, Keisuke Suzuki, Hiroyuki Iizuka
  • A special session in ALIFE2018: Hybrid Life: Approaches to integrate biological, artificial and cognitive systems
    23 Jul. 2018 - 27 Jul. 2018
    Planning etc, Panel chair etc, Peer review
    Academic society etc
    Manuel Baltieri, Keisuke Suzuki, Hiroyuki Iizuka
  • Time in Tokyo : International Symposium on temporal perception and experience
    11 Oct. 2016 - 12 Oct. 2016
    Planning etc, Panel chair etc, Peer review
    Competition etc
    Keisuke Suzuki;Warrick Roseboom;Ryota Kanai;Shigeru Kitazawa;Takashi Ikegami
Social Contribution Activities
  • 人工知能で幻覚体験は再現できるか?
    15 May 2026 - 15 May 2026
    Lecturer
    NPO法人かわさき市民アカデミー事務局
    2026年度前期かわさき市民アカデミー 科学が拓く新世界「人間の知能、人工の知能」講座
    51596714
Media Coverage
  • People who are easily distracted by smartphones are more physiologically reactive, less attuned to their bodies: Study
    08 Apr. 2025
    medicalXpress
    https://medicalxpress.com/news/2025-04-people-easily-distracted-smartphones-physiologically.html, 33902486, [Internet]
  • Scientists Alter Consciousness Without Drugs Using 'Hallucination Machine'
    Nov. 2017
    Yahoo! News
    https://www.yahoo.com/news/scientists-alter-consciousness-without-drugs-203900606.html, [Paper]
  • Hallucination Machine Gives Users Experience Of Trip Without Drugs
    Nov. 2017
    Newsweek
    https://www.newsweek.com/hallucination-machine-gives-users-experience-trip-without-drugs-722607, [Paper]
  • The ‘hallucination machine’ that uses virtual reality and AI to send your brain on a drug-free psychedelic trip
    Nov. 2017
    Other than myself
    Daily Mail
    https://www.dailymail.co.uk/sciencetech/article-5124517/Hallucination-machine-gives-drug-free-psychedelic-trip.html, [Paper]
  • Trip without the high on Sussex University’s hallucination machine
    Nov. 2017
    Other than myself
    The Times
    https://www.thetimes.co.uk/article/trip-without-the-high-on-sussex-university-s-hallucination-machine-809spptbl, [Paper]
  • Inception helmet creates alternative reality Substitutional Reality: system could be used to study cognitive dysfunction in psychiatric patients
    Aug. 2012
    Other than myself
    The Guardian
    https://www.theguardian.com/science/neurophilosophy/2012/aug/26/inception-helmet-alternative-reality, [Paper]