理学博士, 早稲田大学, Mar. 2014

複雑系

自己組織化

創発

力学系

統計力学

カオス

ゆらぎ

エルゴード問題

地震統計

生物物理

機械学習

有限要素法

グラフィックス科学

花弁

微小管

ネマチックテンソル解析

反応拡散

チューリングパターン

重力屈性

根端形状

樹形

点群データ解析

枝分かれ構造

原子間力顕微鏡

胚発生

イネの倒伏

らせん

ハエトリソウ

魚の遊泳

ラグランジュ協同構造

Natural Science, Mathematical physics and fundamental theory of condensed matter physics, 統計力学

Natural Science, Biophysics, chemical physics and soft matter physics

Life Science, Developmental biology

Life Science, Morphology and anatomical structure

Manufacturing Technology (Mechanical Engineering, Electrical and Electronic Engineering, Chemical Engineering), Mechanics of materials and materials

Manufacturing Technology (Mechanical Engineering, Electrical and Electronic Engineering, Chemical Engineering), Fluid engineering

Environmental Science/Agriculture Science, Crop production science

Nanotechnology/Materials, Bio chemistry

Deformation Gradient Tensor Model of Roll–Spiral Transformation for Protein Assembly R-Body
Satoru Tsugawa; Kosuke Kikuchi; Koki Date; Tomonobu Nonoyama; Zichen Kang; Takafumi Ueno
30 May 2026

Size–curvature constraint in the closing motion of Venus flytrap leaves
Michiko Hirata; Zichen Kang; Hiroki Asakawa; Hiraku Suda; Masatsugu Toyota; Yuji Ohashi; Satoru Tsugawa
PLOS One, 26 May 2026
Scientific journal

Temporal changes in surface tension guide microtubule organization and accurate asymmetric division of Arabidopsis zygotes
Zichen Kang; Sakumi Nakagawa; Hikari Matsumoto; Yukitaka Ishimoto; Tomonobu Nonoyama; Yuga Hanaki; Satoru Tsugawa; Minako Ueda
Nature Communications, 22 May 2026
Scientific journal

Ca²⁺ oscillations promote microtubule-band turnover and support tip growth in Arabidopsis zygotes
Hikari Matsumoto; Zichen Kang; Tomonobu Nonoyama; Yusuke Kimata; Satoru Tsugawa; Minako Ueda
20 Feb. 2026

Two-step polar plastid migration via F-actin and microtubules ensures unequal inheritance during asymmetric division of Arabidopsis zygote
Keigo Tada; Hikari Matsumoto; Takao Oi; Zichen Kang; Tomonobu Nonoyama; Satoru Tsugawa; Yusuke Kimata; Shuhei Kusano; Shinya Hagihara; Shintaro Ichikawa; Yutaka Kodama; Minako Ueda
20 Jan. 2026

KymoTip: high‐throughput characterization of tip‐growth dynamics in plant cells
Zichen Kang; Yusuke Kimata; Tomonobu Nonoyama; Toru Ikeuchi; Kazuyuki Kuchitsu; Satoru Tsugawa; Minako Ueda
The Plant Journal, Jan. 2026, [Peer-reviewed]
Scientific journal

Plausible Driving Forces in Elliptic Droplet Self-propulsion
Yuhei Inoue; Tomonobu Nonoyama; Zichen Kang; Shunsuke Yabunaka; Satoru Tsugawa
Journal of the Physical Society of Japan, 15 Sep. 2025, [Peer-reviewed], [Last author, Corresponding author]
Scientific journal

Inference of mechanical forces through 3D reconstruction of the closing motion in venus flytrap leaves
Satoru Tsugawa; Hiroki Asakawa; Michiko Hirata; Tomonobu Nonoyama; Zichen Kang; Masatsugu Toyota; Hiraku Suda
Scientific Reports, 15, 1, Springer Science and Business Media LLC, 28 Jul. 2025, [Peer-reviewed], [Lead author, Corresponding author]
Scientific journal

Simultaneous quantification of cell wall elasticity and turgor pressure in live plant cells by elastic shell theory analysis and AFM
Yuki Yamasaki; Kazunori Okano; Tetsuro Mimura; Satoru Tsugawa; Yoichiroh Hosokawa
Planta, 261, 6, Springer Science and Business Media LLC, 08 May 2025, [Peer-reviewed], [Corresponding author]
Scientific journal

Agent-Based Simulation of Cortical Microtubule Band Movement in Arabidopsis Zygotes
Tomonobu Nonoyama; Zichen Kang; Hikari Matsumoto; Sakumi Nakagawa; Minako Ueda; Satoru Tsugawa
18 Oct. 2024, [Peer-reviewed], [Last author, Corresponding author]

A viscoelastic–plastic deformation model of hemisphere-like tip growth in Arabidopsis zygotes
Zichen Kang; Tomonobu Nonoyama; Yukitaka Ishimoto; Hikari Matsumoto; Sakumi Nakagawa; Minako Ueda; Satoru Tsugawa
Quantitative Plant Biology, 19 Jul. 2024, [Peer-reviewed], [Last author, Corresponding author]
Scientific journal

Shoot gravitropism and organ straightening cooperate to arrive at a mechanically favorable shape in Arabidopsis.
Satoru Tsugawa; Yuzuki Miyake; Keishi Okamoto; Masatsugu Toyota; Hiroki Yagi; Miyo Terao Morita; Ikuko Hara-Nishimura; Taku Demura; Haruko Ueda
Scientific reports, 13, 1, 11165, 11165, 17 Jul. 2023, [Peer-reviewed], [Lead author, Corresponding author], [International Magazine]
English, Scientific journal, Gravitropism is the plant organ bending in response to gravity, while a straightening mechanism prevents bending beyond the gravitropic set-point angle. The promotion and prevention of bending occur simultaneously around the inflorescence stem tip. How these two opposing forces work together and what part of the stem they affect are unknown. To understand the mechanical forces involved, we rotated wild type and organ-straightening-deficient mutant (myosin xif xik) Arabidopsis plants to a horizontal position to initiate bending. The mutant stems started to bend before the wild-type stems, which led us to hypothesize that the force preventing bending was weaker in mutant. We modeled the wild-type and mutant stems as elastic rods, and evaluated two parameters: an organ-angle-dependent gravitropic-responsive parameter (β) and an organ-curvature-dependent proprioceptive-responsive parameter (γ). Our model showed that these two parameters were lower in mutant than in wild type, implying that, unexpectedly, both promotion and prevention of bending are weak in mutant. Subsequently, finite element method simulations revealed that the compressive stress in the middle of the stem was significantly lower in wild type than in mutant. The results of this study show that myosin-XIk-and-XIf-dependent organ straightening adjusts the stress distribution to achieve a mechanically favorable shape.

Thickness-stiffness trade-off improves lodging resistance in rice.
Satoru Tsugawa; Hiroyuki Shima; Yukitaka Ishimoto; Kazuya Ishikawa
Scientific reports, 13, 1, 10828, 10828, 04 Jul. 2023, [Peer-reviewed], [Lead author, Corresponding author], [International Magazine]
English, Scientific journal, Lodging of cereal crops significantly reduces grain yield and quality, making lodging resistance a prime target for breeding programs. However, lodging resistance among different rice (Oryza sativa L.) cultivars in the field remains largely unknown, as is the relationship between the major properties of culms such as their morphological and mechanical properties. Here, we investigated the morphological and mechanical properties of 12 rice cultivars by considering different internodes within culms. We detected variation in these two traits among cultivars: one set of cultivars had thicker but softer culms (thickness-type), while the other set of cultivars showed stiffer but thinner culms (stiffness-type). We designate this variation as a thickness-stiffness trade-off. We then constructed a mechanical model to dissect the mechanical and/or morphological constraints of rice culms subjected to their own weight (self-weight load). Through modeling, we discovered that ear weight and the morphology of the highest internode were important for reducing deflection, which may be important factors to achieve higher lodging resistance. The mechanical theory devised in this study could be used to predict the deflection of rice culms and may open new avenues for novel mechanics-based breeding techniques.

Pulvinar slits: Cellulose-deficient and de-methyl-esterified pectin-rich structures in a legume motor cell
Masahiro Takahara; Satoru Tsugawa; Shingo Sakamoto; Taku Demura; Miyuki T Nakata
Plant Physiology, kiad105, 02 Jun. 2023, [Peer-reviewed]
Scientific journal

A mechanical theory of competition between plant root growth and soil pressure reveals a potential mechanism of root penetration
Haruka Tomobe; Satoru Tsugawa; Yuki Yoshida; Tetsuya Arita; Allen Yi-Lun Tsai; Minoru Kubo; Taku Demura; Shinichiro Sawa
Scientific Reports, 13, 1, 7473, Springer Science and Business Media LLC, 09 May 2023, [Peer-reviewed], [Lead author, Corresponding author]
Scientific journal, Abstract

Root penetration into the soil is essential for plants to access water and nutrients, as well as to mechanically support aboveground structures. This requires a combination of healthy plant growth, adequate soil mechanical properties, and compatible plant–soil interactions. Despite the current knowledge of the static rheology driving the interactions at the root–soil interface, few theoretical approaches have attempted to describe root penetration with dynamic rheology. In this work, we experimentally showed that radish roots in contact with soil of specific density during a specific growth stage fail to penetrate the soil. To explore the mechanism of root penetration into the soil, we constructed a theoretical model to explore the relevant conditions amenable to root entry into the soil. The theory indicates that dimensionless parameters such as root growth anisotropy, static root–soil competition, and dynamic root–soil competition are important for root penetration. The consequent theoretical expectations were supported by finite element analysis, and a potential mechanism of root penetration into the soil is discussed.

Coordinate Normalization of Live-Cell Imaging Data Reveals Growth Dynamics of the Arabidopsis Zygote
Zichen Kang; Hikari Matsumoto; Tomonobu Nonoyama; Sakumi Nakagawa; Yukitaka Ishimoto; Satoru Tsugawa; Minako Ueda
Plant and Cell Physiology, Oxford University Press (OUP), 20 Mar. 2023, [Peer-reviewed], [Corresponding author]
Scientific journal, Abstract

Polarization of the zygote defines the body axis during plant development. In Arabidopsis (Arabidopsis thaliana), the zygote becomes polarized and elongates in the longitudinal direction, ultimately forming the apical–basal axis of the mature plant. Despite its importance, the mechanism for this elongation remains poorly understood. Based on live-cell imaging of the zygote, we developed new image analysis methods, referred to as coordinate normalization, that appropriately fix and align positions in an image, preventing fluctuation across a temporal sequence of images. Using these methods, we discovered that the zygote elongates only at its apical tip region, similar to tip-growing cells such as pollen tubes and root hairs. We also investigated the spatiotemporal dynamics of the apical tip contour of the zygote and observed that the zygote tip retains its isotropic, hemispherical apical shape during cell elongation. By looking at the elliptical fitting of the contour over time, we further discovered that the apical cell tip becomes thinner at first and then thickens, with a transient increase in growth speed that is followed by the first cell division. We performed the same series of analyses using root hairs and established that the hemispherical tip shape and the changes in growth rate associated with changes in tip size are both specific to the zygote. In summary, the Arabidopsis zygote undergoes directional elongation as a tip-growing cell, but its tip retains an unusual isotropic shape, and the manner of growth changes with the developmental stage.

Rigidity Control Mechanism by Turgor Pressure in Plants
Tohya Kanahama; Satoru Tsugawa; Motohiro Sato
Scientific Reports, 13, 2063, 04 Feb. 2023, [Peer-reviewed]
Scientific journal

Femtosecond laser perforation of plant cells for evaluation of cell stiffness
Yuki Yamasaki; Satoru Tsugawa; Kenshiro Ito; Kazunori Okano; Yoichiroh Hosokawa
Applied Physics A, 30 Aug. 2022, [Peer-reviewed]

Elastic shell theory for plant cell wall stiffness reveals contributions of cell wall elasticity and turgor pressure in AFM measurement
Satoru Tsugawa; Yuki Yamasaki; Shota Horiguchi; Tianhao Zhang; Takara Muto; Yosuke Nakaso; Kenshiro Ito; Ryu Takebayashi; Kazunori Okano; Eri Akita; Ryohei Yasukuni; Taku Demura; Tetsuro Mimura; Ken'ichi Kawaguchi; Yoichiroh Hosokawa
Scientific Reports, 12, 13044, 01 Aug. 2022, [Peer-reviewed], [Lead author, Corresponding author]
Scientific journal

Exploring the mechanical and morphological rationality of tree branch structure based on 3D point cloud analysis and the finite element method
Satoru Tsugawa; Kaname Teratsuji; Fumio Okura; Koji Noshita; Masaki Tateno; Jingyao Zhang; Taku Demura
Scientific reports, 12, 1, 4054, 4054, 08 Mar. 2022, [Peer-reviewed], [Lead author, Corresponding author], [International Magazine]
English, Scientific journal

Reduction in organ-organ friction is critical for corolla elongation in morning glory
Ayaka Shimoki; Satoru Tsugawa; Keiichiro Ohashi; Masahito Toda; Akiteru Maeno; Tomoaki Sakamoto; Seisuke Kimura; Takashi Nobusawa; Mika Nagao; Eiji Nitasaka; Taku Demura; Kiyotaka Okada; Seiji Takeda
Communications biology, 4, 1, 285, 285, 05 Mar. 2021, [Peer-reviewed], [Lead author], [International Magazine]
English, Scientific journal

Tissue growth constrains root organ outlines into an isometrically scalable shape
Motohiro Fujiwara; Tatsuaki Goh; Satoru Tsugawa; Keiji Nakajima; Hidehiro Fukaki; Koichi Fujimoto
Development (Cambridge, England), 148, 4, 26 Feb. 2021, [Peer-reviewed], [International Magazine]
English, Scientific journal

Root shape adaptation to mechanical stress derived from unidirectional vibrations in Populus nigra
Marcel Pascal Beier; Satoru Tsugawa; Taku Demura; Toru Fujiwara
Plant biotechnology (Tokyo, Japan), 37, 4, 423, 428, 25 Dec. 2020, [Peer-reviewed], [Domestic magazines]
English, Scientific journal

Spatio-temporal kinematic analysis of shoot gravitropism in Arabidopsis thaliana
Satoru Tsugawa; Norihiro Kanda; Moritaka Nakamura; Tatsuaki Goh; Misato Ohtani; Taku Demura
Plant biotechnology (Tokyo, Japan), 37, 4, 443, 450, 25 Dec. 2020, [Peer-reviewed], [Lead author, Corresponding author], [Domestic magazines]
English, Scientific journal

Laser micromarking technique in studying the negative gravitropism in pea stem
Akira Watanabe; Ashiqur Rahman; Kazuhiko Nishitani; Rie Yamada; Mariko Takahashi; Satoru Tsugawa; Taku Demura
Plant biotechnology (Tokyo, Japan), 37, 4, 485, 488, 25 Dec. 2020, [Peer-reviewed], [Domestic magazines]
English, Scientific journal

Cortical tension overrides geometrical cues to orient microtubules in confined protoplasts
Leia Colin; Antoine Chevallier; Satoru Tsugawa; Florian Gacon; Christophe Godin; Virgile Viasnoff; Timothy E Saunders; Olivier Hamant
Proceedings of the National Academy of Sciences of the United States of America, 117, 51, 32731, 32738, 22 Dec. 2020, [Peer-reviewed], [International Magazine]
English, Scientific journal

A mathematical model explores the contributions of bending and stretching forces to shoot gravitropism in Arabidopsis
Satoru Tsugawa; Tomohiko G. Sano; Hiroyuki Shima; Miyo Terao Morita; Taku Demura
Quantitative Plant Biology, 1, Cambridge University Press ({CUP}), 15 Dec. 2020, [Peer-reviewed], [Lead author, Corresponding author]
Scientific journal

Role of unstable periodic orbits in bubbling weak generalized synchronization
Noi Takahashi; Satoru Tsugawa
PHYSICA D-NONLINEAR PHENOMENA, 414, Dec. 2020, [Peer-reviewed], [Last author, Corresponding author]
English, Scientific journal

Robust organ size requires robust timing of initiation orchestrated by focused auxin and cytokinin signalling
Mingyuan Zhu; Weiwei Chen; Vincent Mirabet; Lilan Hong; Simone Bovio; Soeren Strauss; Erich M Schwarz; Satoru Tsugawa; Zhou Wang; Richard S Smith; Chun-Biu Li; Olivier Hamant; Arezki Boudaoud; Adrienne H K Roeder
Nature plants, 6, 6, 686, 698, Jun. 2020, [Peer-reviewed], [International Magazine]
English, Scientific journal

Suppression of soft spots and excited modes in the shape deformation model with spatio-temporal growth noise
Satoru Tsugawa
Journal of theoretical biology, 486, 110092, 110092, 07 Feb. 2020, [Peer-reviewed], [Lead author, Last author, Corresponding author], [International Magazine]
English, Scientific journal

A Rho-based reaction-diffusion system governs cell wall patterning in metaxylem vessels
Yoshinobu Nagashima; Satoru Tsugawa; Atsushi Mochizuki; Takema Sasaki; Hiroo Fukuda; Yoshihisa Oda
Scientific reports, 8, 1, 11542, 11542, 01 Aug. 2018, [Peer-reviewed], [International Magazine]
English, Scientific journal

Heterogeneity and Robustness in Plant Morphogenesis: From Cells to Organs
Lilan Hong; Mathilde Dumond; Mingyuan Zhu; Satoru Tsugawa; Chun-Biu Li; Arezki Boudaoud; Olivier Hamant; Adrienne H K Roeder
Annual review of plant biology, 69, 469, 495, 29 Apr. 2018, [Peer-reviewed], [International Magazine]
English

Clones of cells switch from reduction to enhancement of size variability in Arabidopsis sepals
Satoru Tsugawa; Nathan Hervieux; Daniel Kierzkowski; Anne-Lise Routier-Kierzkowska; Aleksandra Sapala; Olivier Hamant; Richard S Smith; Adrienne H K Roeder; Arezki Boudaoud; Chun-Biu Li
Development (Cambridge, England), 144, 23, 4398, 4405, 01 Dec. 2017, [Peer-reviewed], [Lead author], [International Magazine]
English, Scientific journal

Mechanical Shielding of Rapidly Growing Cells Buffers Growth Heterogeneity and Contributes to Organ Shape Reproducibility
Nathan Hervieux; Satoru Tsugawa; Antoine Fruleux; Mathilde Dumond; Anne-Lise Routier-Kierzkowska; Tamiki Komatsuzaki; Arezki Boudaoud; John C Larkin; Richard S Smith; Chun-Biu Li; Olivier Hamant
Current biology : CB, 27, 22, 3468, 3479, 20 Nov. 2017, [Peer-reviewed], [Lead author], [International Magazine]
English, Scientific journal

Variable Cell Growth Yields Reproducible OrganDevelopment through Spatiotemporal Averaging
Lilan Hong; Mathilde Dumond; Satoru Tsugawa; Aleksandra Sapala; Anne-Lise Routier-Kierzkowska; Yong Zhou; Catherine Chen; Annamaria Kiss; Mingyuan Zhu; Olivier Hamant; Richard S Smith; Tamiki Komatsuzaki; Chun-Biu Li; Arezki Boudaoud; Adrienne H K Roeder
Developmental cell, 38, 1, 15, 32, 11 Jul. 2016, [Peer-reviewed], [Lead author], [International Magazine]
English, Scientific journal

Extracting Subcellular Fibrillar Alignment with Error Estimation: Application to Microtubules
Satoru Tsugawa; Nathan Hervieux; Oliver Hamant; Arezki Boudaoud; Richard S Smith; Chun-Biu Li; Tamiki Komatsuzaki
Biophysical journal, 110, 8, 1836, 1844, 26 Apr. 2016, [Peer-reviewed], [Lead author], [International Magazine]
English, Scientific journal

Statistical Laws of the Ensemble Average Associated with Allan Variance: Special Issue on New Challenges in Complex Systems Science
Satoru Tsugawa; Yoji Aizawa
理工研報告特集号 : ASTE : advances in science, technology and environmentology : special issue / 早稲田大学理工学研究所ASTE70周年特集号編集委員会 編, 11, 197, 199, Mar. 2015, [Peer-reviewed], [Lead author]
English

Stagnant Motion in Hamiltonian Dynamics -Mushroom Billiard Case with Smooth Outermost KAM Tori-
Satoru Tsugawa; Yoji Aizawa
JOURNAL OF THE PHYSICAL SOCIETY OF JAPAN, 83, 2, Feb. 2014, [Peer-reviewed], [Lead author]
English, Scientific journal

Aftershock Cascade of the 3.11 Earthquake (2011) in Fukushima-Miyagi Area
Yoji Aizawa; Satoru Tsugawa
NONLINEAR PHENOMENA IN COMPLEX SYSTEMS: FROM NANO TO MACRO SCALE, 21, 33, 2014, [Peer-reviewed]
English, International conference proceedings

Seismic statistics: Universality and interim report on the 3. 11 earthquake (2011) in Fukushima-Miyagi area
Yoji Aizawa; Tomohiro Hasumi; Satoru Tsugawa
Nonlinear Phenomena in Complex Systems, 16, 2, 116, 130, 2013, [Peer-reviewed]
Scientific journal

Stagnant Motion in Chaotic Region of Mushroom Billiard System with Dielectric Medium
Satoru Tsugawa; Yoji Aizawa
JOURNAL OF THE PHYSICAL SOCIETY OF JAPAN, 81, 6, Jun. 2012, [Peer-reviewed], [Lead author]
English, Scientific journal

Analysis of the Mechanism of Cortical Microtubule Reorganization at the M/G1 interface
小野華歌; 野々山朋信; KANG Zichen; 植田美那子; 津川暁; 檜垣匠, バイオイメージング, 33, 2, 2024

M/G1境界期にトランスバースに配向を確立する表層微小管の画像解析
小野華歌; 野々山朋信; 康子辰; 植田美那子; 津川暁; 檜垣匠, 日本植物学会大会研究発表記録(CD-ROM), 88th, 2024

Analysis of a mathematical model of shoot gravitropism in Arabidopsis thaliana
Tsugawa Satoru; Sano Tomohiko G.; Shima Hiroyuki; Morita Miyo Terao; Demura Taku, PLANT MORPHOLOGY, 33, 1, 71, 76, 08 Apr. 2022, [Peer-reviewed], [Invited], [Lead author]
Plant shoot gravitropism is determined by gravity sensing, curvature sensing (proprioception), and the ability to uphold self-weight. Recently, data- and model-analyses have revealed the detailed morphology of shoot bending, while mechanical mechanism to cause the bending is still unknown. By making a best correspondence between data and model of shoot bending in wild-type and lazy1-like 1(lzy1) mutant Arabidopsis thaliana, we found that both the bending force (derived from the gravi-proprioceptive response) and the stretching force (derived from shoot axial growth) differ significantly between the wild type and mutant. Finally, we interpreted the mechanical forces associated with differential cell growth and provide a plausible mechanical explanation of shoot gravitropism., The Japanese Society of Plant Morphology, Japanese

A Physical Model to Identify the Common Organ Shape Across Species
Tatsuaki GOH; Motohiro FUJIWARA; Satoru TSUGAWA; Koichi FUJIMOTO, Seibutsu Butsuri, 62, 1, 7, 12, 2022, [Peer-reviewed], [Invited]
Organ shape is similar across species despite their size diversity. Here we review that the physical model of a mathematical formula representing the organ shape can identify a developmental mechanism ensuring the similarity. We recently found that the shape of plant root organ outlines can be commonly described as a catenary curve, a stable structure found in a free-hanging chain. The physical model of catenary curve predicts that the tissue growth rules force to form the catenary-curved root tip. The physical model-inspired research strategy provides new insights for the shared constraints on the shape similarity., Biophysical Society of Japan, Japanese

がく片器官の形の頑健性：細胞成長揺らぎの時空間平均化の仕組み
津川 暁, 植物科学の最前線, 10, A, 22, 31, 2019, [Peer-reviewed], [Invited], [Lead author]
公益社団法人 日本植物学会

Mechanism of Plant Morphogenesis Approached by Data Science
TSUGAWA Satoru, Seibutsu Butsuri, 57, 3, 131, 134, 2017, [Peer-reviewed], [Invited], [Lead author]
How do organs reach their reproducible shapes? This question remains open because it has been difficult to connect the microscopic dynamics/stochasticity to the macroscopic regulation/reproducibility at the molecular-cellular-organ level. We challenged by data science to quantify the heterogeneity of microtubule alignments to connect the molecular dynamics to the regulation of cell shape, and to quantify the variability of cell growth to connect the cellular stochasticity to the reproducible organ shape. Our key and counterintuitive result was “cellular randomness is required for reaching the correct organ size and shape” meaning that the order can be emerged from disorder., The Biophysical Society of Japan General Incorporated Association, Japanese

理論生物学事典
巌佐, 庸; 大槻, 久; 佐竹, 暁子; 望月, 敦史, 細胞からの組織の秩序形成
朝倉書店, Nov. 2025, 9784254172034, xiv, 548p, 図版4p(ページ付なし), Japanese, [Contributor]

AFM計測と弾性シェル理論による植物細胞の膨圧・細胞壁硬さの同時推定
津川暁
2025年度日本顕微鏡学会SPM研究会, Invited oral presentation
Dec. 2025, [Invited]

植物細胞の集団的協調現象：植物はどのように成長・屈曲するのか？
津川暁
振動と同期学の研究会, Nov. 2025, Invited oral presentation
[Invited]

Temporal changes in surface tension guide the accurate asymmetric division of Arabidopsis zygotes
S. Tsugawa; Z. Kang; S. Nakagawa; H. Matsumoto; Y. Ishimoto; T. Nonoyama; Y. Hanaki; M. Ueda
The Japanese Society of Plant Physiologists, Mar. 2025, Invited oral presentation
[Invited]

R-bodyのコイル-らせん転移における変形設計原理について
津川暁
第4会発動分子科学研究会, Sep. 2023, Invited oral presentation
[Invited]

植物の構造と機能に学ぶバイオミメティックス
津川暁
分子研研究会「生体分子材料を探る：発動分子のさらなる理解と設計に向けて」, Mar. 2023, Invited oral presentation
[Invited]

応力均一化を基盤とする樹木の成長応力解析
津川暁
日本材料学会木質材料部門委員会第311定例研究会, Jan. 2023, Invited oral presentation
[Invited]

植物の「カタチ」と「チカラ」を解き明かす
津川暁
MIMS研究集会, Nov. 2021
[Invited]

応力均一化仮説に基づく樹木の屈曲解析とその建築構造物への応用可能性
津川 暁; 出村 拓; 細見 亮太; 小御門 真伍
「シェル・空間構造」セミナー, Dec. 2020, Invited oral presentation
[Invited]

植物の形と動きの定量化の先のモデル化の試み
津川暁
MIMS研究集会, Nov. 2020, Invited oral presentation
[Invited]

シロイヌナズナ花茎重力屈性の数理モデル解析
津川 暁; 佐野友彦; 島弘幸; 森田(寺尾)美代; 出村拓
第84回植物学会, Sep. 2020, Invited oral presentation
[Invited]

A mathematical model explores the contributions of bending and stretching forces to shoot gravitropism in Arabidopsis
Satoru Tsugawa
Quantitative Plant Biology Webinar, Apr. 2020, Invited oral presentation
[Invited]

MorphoGraphXによる３Dイメージングデータの定量的解析
津川暁
光塾,, Dec. 2019
[Invited]

萼片器官の形の頑健性：細胞成長揺らぎの時空間平均化の仕組み
津川暁
第82回日本植物学会, Sep. 2018, Invited oral presentation
[Invited]

Organ shape robustness through cell variability
津川暁
Biological Symposium, Aug. 2018, Invited oral presentation
[Invited]

植物細胞の確率的振る舞いと器官形状の頑健性
津川暁
散逸構造・カオス・複雑系シンポジウム~イリヤ.プリゴジン先生の業績を偲んで~, May 2018, Invited oral presentation
[Invited]

植物器官の形の頑健性：力学的弱点を時空間的に緩和するメカニズム
津川暁
自然知能研究グループ（NI）-複雑コミュニケーション科学研究会(CCS)合同研究会, Mar. 2018, Invited oral presentation
[Invited]

多細胞間での時空間的相互作用の理解を目指した定量的解析基盤の創出
CREST
Oct. 2021 - Mar. 2027
国立研究開発法人科学技術振興機構

Coincident estimation of cell elasticity and intracellular pressure by atomic force measurement and its elastic shell theory analysis
Grants-in-Aid for Scientific Research
01 Apr. 2023 - 31 Mar. 2026
細川 陽一郎; 津川 暁; 丹賀 直美
Japan Society for the Promotion of Science, Grant-in-Aid for Scientific Research (B), Nara Institute of Science and Technology, 23K25840

植物重力屈性の力学的仕組みの解明
科学研究費助成事業 若手研究
01 Apr. 2020 - 31 Mar. 2024
津川 暁
今年度は，シロイヌナズナ花茎の重力屈性挙動について研究し，その成果を論文として発表した．今年度の助成金は，研究成果論文の雑誌掲載費と文献購入費に充てた．


1.一報目の論文はPlant Biotechnology誌の特集号に掲載された．本論文では，野生型シロイヌナズナ花茎の重力屈性を理解するために，シロイヌナズナ花茎が水平に倒れた状態から上方に起き上がる挙動をタイムラプス動画で撮影した．得られた動画に対して画像データ解析を適用し，花茎のどの部分がいつ曲がり始めるかを定量的に分析したところ，約50分後に花茎先端付近から曲がり始め，約100分後には花茎中心部に曲がり位置が移動することを確認できた．この定性的な観測を屈曲に異常がある変異体と比較するために，野生型において曲がり始め時間や曲がり位置を定量的に特徴づける方法を提案している．また，実際の屈曲挙動を模倣する1次元弾性体による数理モデルを構築し，植物の屈曲挙動を力学的に理解する方法論を確立した．
2.二報目の論文はQuantitative Plant Biology誌に掲載された．本論文では，上記で得られた画像データ分析手法を適用し，野生型シロイヌナズナと屈曲が遅くなる変異体の屈曲挙動の違いを明らかにしている．特に花茎の曲がり具合を示す最大曲率が野生型の方が明確に高くなることを定量的に示した，また，本研究で画像データ結果と数理モデル結果を照合・比較する従来にない研究手法を確立し，花茎のどの部分がいつ曲がるかを力学的に検証する基礎的な方法を確立した．分析の結果，野生型花茎が変異体花茎に比べてより強い偏差成長を発揮することで，変異体よりも早く花茎を曲げていることを明らかにした．
3.他二報の論文は，上記研究を発展させたアサガオ花弁の座屈の研究と花茎屈曲を定量的に分析する実験改良の研究であり，今後の学術連携促進が期待される．
日本学術振興会, 若手研究, 奈良先端科学技術大学院大学, 20K15832

Characteristics of weak chaos in view of large deviation analysis and infinite ergodicity
Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research (C)
01 Apr. 2013 - 31 Mar. 2017
Aizawa Yoji; TSUGAWA Satoru; NAKAGAWA Masaki
There still remain many unsolved problems in the nonstationary regime of chaotic or turbulent phenomena; for instance, anomalous diffusion in hamiltonian systems, intermittency, and infinite ergodicity, et cetera. In this research, we studied several nonlinear dynamical models, such as the hamiltonian dynamics of lattice vibrations, mushroom billiards with clear boundaries between chaos and torus, antlion models with dissipative infinite measure ergodicity, nonchaotic strange attractors in periodically driven systems, wave turbulence in Bloch equation, etc., and succeeded to elucidate the diversity of the nonstationary chaos and its universal aspects in statistical behaviors. Furthermore, we studied a new method for the numerical analysis of chaotic intermittent time series by using the real data of interoccurrence time statistics of earthquakes, and succeeded to obtain some new statistical laws of magnitude correlations in the shock sequences.
Japan Society for the Promotion of Science, Grant-in-Aid for Scientific Research (C), Waseda University, 25400411